Environment: milieu / climate zone / depth range / distribution range					
						Ecology					
				
				
				
					Marine;  demersal; oceanodromous (Ref. 51243); depth range 5 - 50 m (Ref. 6302). Temperate; 64°N -   26°N, 24°W -   42°E				
				 
			
			
			
				
				
				
					Eastern Atlantic:  68°N to Morocco.  Reported from Iceland (Ref. 12462). Also known throughout the Mediterranean and Black Sea.
				
				 
			
			
				
					Length at first maturity / Size / Weight / Age
				
				
				
					Maturity: Lm 37.0, range 33 -  41 cm
 Max length : 75.0 cm TL male/unsexed; (Ref. 35388); common length : 30.0 cm TL male/unsexed; (Ref. 3397); max. published weight: 8.0 kg (Ref. 35388); max. reported age: 6 years (Ref. 38118)				
				 
			
			
						
				
				
				
					Thinner and more slender body.  Skin with small, smooth scales, without bony tubercle.  Able to change color of the eyed side, matching the bottom they rest on.  Color is often olive green, with dark and light spots (Ref. 35388).
Body shape (shape guide): short and / or deep.				
				 
			
			
			
			
			
			
			
				Lives on sandy or mixed bottoms; feeds on bottom-living fishes and larger crustaceans.  Marketed fresh and frozen; eaten steamed, fried, broiled, boiled, microwaved and baked (Ref. 9988).			
			 
			
			
			
				
				
				
								
				
			
			 
				
				
				
					Bauchot, M.-L., 1987. Poissons osseux. p. 891-1421. In W. Fischer, M.L. Bauchot and M. Schneider (eds.) Fiches FAO d'identification pour les besoins de la pêche. (rev. 1). Méditerranée et mer Noire. Zone de pêche 37. Vol. II. Commission des Communautés Européennes and FAO, Rome. (Ref. 3397)
				
				 
			
			
			
							
					
						IUCN Red List Status   (Ref. 130435: Version 2025-1)
					
					
				 
					
			
			
			
			
			
			
			
				
					Threat to humans  
				
				
				
					  Harmless				
				
			 
			
			
			
			
				
					Human uses  				
				
				
					Fisheries: commercial; gamefish: yes				
				
				
			
			
						
			
			
			
				
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					Estimates based on models				
				
				
				Preferred temperature (Ref. 
123201): 8.4 - 19.5, mean 11.2 °C (based on 472 cells).
				
				
					
					Phylogenetic diversity index  (Ref. 
82804):  PD
50 = 0.5645   [Uniqueness, from 0.5 = low to 2.0 = high].					
													Bayesian length-weight: a=0.01096 (0.00893 - 0.01346), b=3.01 (2.95 - 3.07), in cm total length, based on LWR estimates for this species (Ref. 
93245).
					
					Trophic level  (Ref. 
69278):  4.4   ±0.1 se; based on diet studies.					
						 Generation time: 4.1 ( na - na) years. Estimated as median ln(3)/K based on 2
 growth studies.					
						Resilience  (Ref. 
120179):  Medium, minimum population doubling time 1.4 - 4.4 years (tm=3-5; tmax=6).					
						 Prior r = 0.47, 95% CL = 0.31 - 0.70, Based on 5 full stock assessments.					
				
									
					Fishing Vulnerability  (Ref. 
59153):  Moderate vulnerability (36 of 100). 
🛈					
					Climate Vulnerability  (Ref. 
125649):  Moderate to high vulnerability (54 of 100). 
🛈					
						Nutrients  (Ref. 
124155):  Calcium = 37.7 [14.9, 70.5] mg/100g; Iron = 0.174 [0.061, 0.770] mg/100g; Protein = 19.3 [17.8, 20.8] %; Omega3 = 0.262 [0.140, 0.491] g/100g; Selenium = 14 [6, 33] μg/100g; VitaminA = 8.74 [1.87, 40.75] μg/100g; Zinc = 0.145 [0.082, 0.500] mg/100g (wet weight); based on 
nutrient studies.