Environment: milieu / climate zone / depth range / distribution range
Marine; demersal; depth range 15 - 400 m (Ref. 26999), usually - m (Ref. ). Subtropical; 0°C - ; 45°N - 15°S, 26°W - 36°E
Eastern Atlantic: Mediterranean and the Adriatic Sea (Ref. 12382); Mauritania to Angola, including Madeira, Cape Verde and the Canary Islands (Ref. 5979).
Size / Weight / Age
Maturity: Lm ?  range ? - ? cm
Max length : 45.0 cm TL male/unsexed; (Ref. 27584); common length : 13.0 cm TL male/unsexed; (Ref. 3397); max. published weight: 700.00 g (Ref. 27584)
soft rays: 63 - 73. The scales on the eyed side are ctenoid and cycloid on the blind side. The inter-orbital width is much wider in males. The upper pectoral fin rays are not prolonged. The subspecies in the Mediterranean and African coast, P. podas has 75-86 lateral line scales, with 13-20 scales making up the curved portion. Its eyed side has alight brown color, bearing darker spots. The subspecies in Madeira and the Canaries, B. p. maderensis, has 88-91 lateral line scales, with 20-21 on the lateral line curve. Its eyed side appears dark brown to dark violet and seldom spotted.
Found in shallow waters, over sandy and muddy bottoms of the continental plateau (Ref. 5377, 12382). Feeds on benthic small fishes and invertebrates. Reproduction occurs between May and August. Small individuals adapt well in aquariums but require sufficient bottom areas (Ref. 12382).
Life cycle and mating behavior
Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae
Aldebert, Y., M. Desoutter and J.-C. Quéro, 1990. Bothidae. p. 1027-1036. In J.C. Quero, J.C. Hureau, C. Karrer, A. Post and L. Saldanha (eds.) Check-list of the fishes of the eastern tropical Atlantic (CLOFETA). JNICT, Lisbon; SEI, Paris; and UNESCO, Paris. Vol. 2. (Ref. 5979)
IUCN Red List Status (Ref. 124695)
Threat to humans
Fisheries: commercial; aquarium: commercial
ReferencesAquacultureAquaculture profileStrainsGeneticsAllele frequenciesHeritabilityDiseasesProcessingNutrientsMass conversion
Estimates based on models
Preferred temperature (Ref. 115969
): 13 - 17.6, mean 15.1 °C (based on 199 cells).
Phylogenetic diversity index (Ref. 82805
= 0.5000 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00871 (0.00648 - 0.01170), b=3.07 (3.01 - 3.13), in cm total length, based on LWR estimates for this species (Ref. 93245
Trophic level (Ref. 69278
): 3.4 ±0.4 se; based on diet studies.
Resilience (Ref. 120179
): Medium, minimum population doubling time 1.4 - 4.4 years (K=0.43).
Vulnerability (Ref. 59153
): High vulnerability (55 of 100) .
Nutrients (Ref. 124155
): Calcium = 70 [36, 156] mg/100g ; Iron = 0.906 [0.555, 1.568] mg/100g ; Protein = 18.5 [17.2, 19.9] % ; Omega3 = 0.231 [0.129, 0.410] g/100g ; Selenium = 28 [14, 59] μg/100g ; VitaminA = 11.7 [3.1, 45.0] μg/100g ; Zinc = 0.698 [0.463, 1.024] mg/100g (wet weight);