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	<dc:identifier>FB-9156</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=9156</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Cichliformes</dwc:Order>
	<dwc:Family>Cichlidae</dwc:Family>
	<dwc:Genus>Tylochromis</dwc:Genus>
	<dwc:ScientificName>Tylochromis variabilis Stiassny, 1989</dwc:ScientificName>
	<commonName xml:lang=''>Ekeki</commonName>
	<commonName xml:lang=''>Ikeki</commonName>
	<commonName xml:lang=''>Loke</commonName>
	<commonName xml:lang=''>Sokoke</commonName>
	<synonym relationship='misapplied name'>Pelmatochromis lateralis (non Boulenger, 1898)</synonym>
	<synonym relationship='misapplied name'>Tylochromis banguelensis (non Regan, 1920)</synonym>
	<synonym relationship='misapplied name'>Tylochromis lateralis (non Boulenger, 1898)</synonym>
	<dcterms:created>1994-03-07</dcterms:created>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=1884&amp;speccode=9156'>Boulenger, G.A.0 Additions à la faune ichthyologique du bassin du Congo. Ann. Mus. Congo, Zool. (1), 2(2):19-57. (Ref. 1884)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=2970&amp;speccode=9156'>Poll, M.0 Exploration du Parc National de l'Upemba - Mission G.F. De Witte en collaboration avec W. Adam, A. Janssens, L. Van Meel et R. Verheyen (1946-1949). Fascicule 73. Poissons. Fondation pour favoriser les Recherches Scientifiques en Afrique, Bruxelles. 127 pp. (Ref. 2970)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4910&amp;speccode=9156'>Gosse, J.P.0 Le milieu aquatique et l'écologie des poissons dans la région de Yangambi. Ann. Mus. R. Afr. Centr., Sér. in-8°, Sci. Zool., 116:113-270. (Ref. 4910)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5714&amp;speccode=9156'>Stiassny, M.L.J.0 Tylochromis. p. 526-533. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 5714)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31982&amp;speccode=9156'>Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41414&amp;speccode=9156'>Anonymous0 Fish collection database of the American Museum of Natural History. American Museum of Natural History, Central Park West , NY 10024-5192, USA. (Ref. 41414)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41590&amp;speccode=9156'>David, L. and M. Poll0 Contribution à la faune ichthyologique du Congo Belge: collections du Dr. H. Schouteden (1924-1926) et d'autres récolteurs. Ann. Mus. Congo Belge, Zool. (1), 3(5):189-294. (Ref. 41590)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41594&amp;speccode=9156'>Nichols, J.T. and L. Griscom0 Fresh-water fishes of the Congo basin obtained by the American Museum Congo Expedition, 1909-1915. Bull. Am. Mus. Nat. Hist. 37(25):653-756. (Ref. 41594)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=42005&amp;speccode=9156'>Fowler, H.W.0 Zoological results of the George Vanderbilt African Expedition of 1934. Part III. The fresh water fishes. Proc. Acad. nat. Sci. Philad. 83:243-335. (Ref. 42005)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=42033&amp;speccode=9156'>Poll, M. and J.P. Gosse0 Contribution à l'étude systématique de la faune ichthyologique du Congo central. Ann. Mus. R. Afr. Centr., Sér. in-8°, Sci. Zool., 116:43-110. (Ref. 42033)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=44840&amp;speccode=9156'>Daget, J. and A. Stauch0 Poissons de la rive droite du Moyen-Congo. Mission A. Stauch (février-avril 1961). Bull. Inst. Rech. Sci. Congo 2:41-48. (Ref. 44840)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=45434&amp;speccode=9156'>Banister, K.E. and R.G. Bailey0 Fishes collected by the Zaïre River Expedition, 1974-75. Zool. J. Linn. Soc. 66:205-249. (Ref. 45434)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=45483&amp;speccode=9156'>Hulot, A.0 Le régime alimentaire des poissons du centre africain. Intérêt éventuel de ces poissons en vue d'une zootechnie économique au Congo belge. Bull. Agric. Congo Belge 41(1):145-176. (Ref. 45483)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=45616&amp;speccode=9156'>Poll, M.0 Contribution à la faune ichthyologique du Katanga. Ann. Mus. Congo Belge, Zool. (1), 3(3):101-152. (Ref. 45616)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=46253&amp;speccode=9156'>Boulenger, G.A.0 On a small collection of fishes from the Kasai River (Congo). Ann. Mag. Nat. Hist. (Ser. 7) 16:640-642. (Ref. 46253)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=48960&amp;speccode=9156'>Depasse, P.0 Monographie piscicole de la province orientale. Description du milieu limnologique; programme et réalisations en matière de pêche et de pisciculture. Bull. Agr. Congo Belge 47(4):959-1088. (Ref. 48960)</reference>
	<reference isbn='3-928819-33-X' url='http://www.fishbase.org/references/FBRefSummary.php?id=52307&amp;speccode=9156'>Lamboj, A.0 The cichlid fishes of Western Africa. Birgit Schmettkamp Verlag, Bornheim, Germany. 255 p. (Ref. 52307)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52345&amp;speccode=9156'>Stiassny, M.L.J.0 &lt;i&gt;Tylochromis&lt;/i&gt;, relationships and the phylogenetic status of the African Cichlidae. Am. Mus. Novit. 2993:1-14. (Ref. 52345)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52346&amp;speccode=9156'>Stiassny, M.L.J.0 A taxonomic revision of the African genus &lt;i&gt;Tylochromis&lt;/i&gt; (Labroidei, Cichlidae); whith notes on the anatomy and relationships of the group. Ann. Mus. R. Afr. Cent. Ser. 8, Sci. Zool., 258:1-161. (Ref. 52346)</reference><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1994-03-07 00:00:00</dcterms:created>
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	<dc:title>Biology</dc:title>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=9156</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>Opportunistic benthic macrophage (Ref. 52346). Feeds on plants, insects and (to a lesser degree) molluscs (Ref. 52307).  Maternal (Ref. 52307) mouthbrooder (Ref. 4910, 52307) that does not pair-bond (Ref. 52307).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52346&amp;speccode=9156'>Stiassny, M.L.J. 1989 A taxonomic revision of the African genus &lt;i&gt;Tylochromis&lt;/i&gt; (Labroidei, Cichlidae); whith notes on the anatomy and relationships of the group. Ann. Mus. R. Afr. Cent. Ser. 8, Sci. Zool., 258:1-161. (Ref. 52346)</reference></dataObject><dataObject>
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	<agent homepage='http://www.africamuseum.be#' logoURL='' role='recorder'>Tobias Musschoot</agent>
	<dcterms:created>2005-03-15 00:00:00</dcterms:created>
	<dcterms:modified>2017-12-11 21:41:37</dcterms:modified>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=9156</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>Diagnosis: 33-35 lateral line scales; 15-16 gill-rakers along lower limb of first arch; entire premaxillary oral valve present; dorsal fin length of 55.4-65.5% SL (Ref. 52346).Description: relatively deep bodied and robust; predorsal profile more or less straight, rising at an angle of about 50-60° to the horizontal; fleshy bulge in interorbital region frequently present; snout length variable: some specimens with a straight-snouted appearance, others with a shorter snout, a somewhat inclined lower jaw and a definitely pugnacious aspect; dorsum variously recurved from nape to caudal peduncle; deepest body depth may be at dorsal fin origin, or as far back as below 5th-6th spine; mouth a little inclined from the horizontal when closed, but inclination rarely strongly marked, except in certain deep bodied, short-snouted individuals from the Luvua River with a quite steeply inclined lower jaw, often at an angle of about 15-20° to the horizontal; lips thick and reasonably fleshy; lower pharyngeal jaw mostly robust and well ossified; inferior vertebral apophysis well developed and robust, and of standard Tylochromis type; broad, crenate gill-rakers closely overlapping for length of lower limb of first arch; 2-3 short hypobranchial rakers usually present; short and thick epibranchial rakers fringe upper limb of first arch; 33-35 lateral line scales (Ref. 52346). 7-8 scales below lower lateral line; 3 scale rows on cheek below eye (Ref. 42005). Upper branch of lateral line usually terminates a little in advance of end of dorsal fin, but may extend full length of fin; terminal canal bearing scales often descend a scale row, canal then continuing for another 2(-3) scales; dorsal and ventral branches of lateral line on caudal fin well developed and extending almost to fin periphery; median branch usually well developed, but rarely extending much beyond 2/3 of caudal fin length; dorsal fin spines increase gradually in length to (5th) 6th-7th and are then more or less equal in length; anal fin rounded; pectoral fins rather short, rarely reaching beyond level of vent; first branched pelvic ray somewhat produced but rarely markedly filamentous; caudal fin slightly emarginate and usually heavily scaled almost to fin periphery (Ref. 52346). Coloration: pearl grey and greenish above in more or less longitudinal stripes, whitish below, silvery troughout; scales of caudal peduncle and flank in larger individuals ringed in dark brown pigment, resulting in a series of longitudinal striations passing along scale rows; longitudinal striping most pronounced on caudal peduncle and dorsum; vertical stripes less prominent in adults and completely absent in specimens of 15cm SL (Ref. 52346). Dorsal fin transparent grey with bright blue reflections on spines and brownish-red longitudinal markings; caudal fin greyish edged with red on lower half (Ref. 52307, 52346). Pectoral fins transparent grey; pelvic and anal fins pale blue-grey (Ref. 52307, 52346), former with red markings anteriorly (Ref. 52346). Anal fin of mature male fish smoky brown with a few rows of pale non-ocellate maculae that are typically lacking in females; mature males exhibit caudal fin striping and spotting; top of head in front of eyes dark grey; gill covers yellowish and an area behind head dusky yellowish grey; some red markings between pelvic and pectoral fins (Ref. 52346). Preserved juveniles: strong nape stripe separated from dark opercular blotch; 7-8 thick vertical strips which extend down onto flank well below midline, often almost to ventrum; juvenile barring usually pronounced in specimens of less than 6-7cm SL; dorsal fin dark grey brown and spotted with numerous rows of non-ocellate pale maculae (Ref. 52346).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52346&amp;speccode=9156'>Stiassny, M.L.J. 1989 A taxonomic revision of the African genus &lt;i&gt;Tylochromis&lt;/i&gt; (Labroidei, Cichlidae); whith notes on the anatomy and relationships of the group. Ann. Mus. R. Afr. Cent. Ser. 8, Sci. Zool., 258:1-161. (Ref. 52346)</reference></dataObject><dataObject>
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	<agent homepage='http://www.africamuseum.be#' logoURL='' role='recorder'>Tobias Musschoot</agent>
	<dcterms:created>2005-03-15 00:00:00</dcterms:created>
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	<dc:description>Dorsal spines (total): 14 - 16; Dorsal soft rays (total): 11 - 16;  Analspines: 3; Analsoft rays: 7 - 8; Vertebrae: 28 - 29</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52346&amp;speccode=9156'>Stiassny, M.L.J. 1989 A taxonomic revision of the African genus &lt;i&gt;Tylochromis&lt;/i&gt; (Labroidei, Cichlidae); whith notes on the anatomy and relationships of the group. Ann. Mus. R. Afr. Cent. Ser. 8, Sci. Zool., 258:1-161. (Ref. 52346)</reference></dataObject><dataObject>
	<dc:identifier>FB-Distribution-9156</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1994-03-07 00:00:00</dcterms:created>
	<dcterms:modified>2016-11-16 21:31:55</dcterms:modified>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=9156</dc:source>
	<dc:description>Africa:  middle and upper Congo River basin, including the Sankuru, Sangha, Ubangui, Uele, Kotto, Bomu, Lomami, Aruwimi, Itimbiri, Lindi-Tshopo, Lualaba, upper Lualaba and Chambezi, in Cameroon, Central African Republic, Democratic Republic of the Congo and Zambia (Ref. 52346). Also reported from the Sanga in Republic of Congo (Ref. 44840).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5714&amp;speccode=9156'>Stiassny, M.L.J. 1991 Tylochromis. p. 526-533. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 5714)</reference></dataObject><dataObject>
	<dc:identifier>FB-TrophicStrategy-9156</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Grace Tolentino Pablico</agent>
	<dcterms:created>2005-09-30 00:00:00</dcterms:created>
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	<audience>General public</audience>
	<dc:source>variabilis</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Opportunistic benthic macrophage (Ref. 52346).  Feeds on plants, insects and (to a lesser degree) molluscs (Ref. 52307).  Mouthbrooder (Ref. 4910).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52346&amp;speccode=9156'>Stiassny, M.L.J. 1989 A taxonomic revision of the African genus &lt;i&gt;Tylochromis&lt;/i&gt; (Labroidei, Cichlidae); whith notes on the anatomy and relationships of the group. Ann. Mus. R. Afr. Cent. Ser. 8, Sci. Zool., 258:1-161. (Ref. 52346)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-9156</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1994-03-07 00:00:00</dcterms:created>
	<dcterms:modified>2024-04-16 00:00:00</dcterms:modified>
	<dc:title>Max. size</dc:title>
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	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=9156</dc:source>
	<dc:description>27.6 cm NG (male/unsexed; (Ref. 2970))</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=2970&amp;speccode=9156'>Poll, M. 1976 Exploration du Parc National de l'Upemba - Mission G.F. De Witte en collaboration avec W. Adam, A. Janssens, L. Van Meel et R. Verheyen (1946-1949). Fascicule 73. Poissons. Fondation pour favoriser les Recherches Scientifiques en Afrique, Bruxelles. 127 pp. (Ref. 2970)</reference></dataObject><dataObject>
	<dc:identifier>FB-Threats-2</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-9156</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1994-03-07 00:00:00</dcterms:created>
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	<dc:title>Environment</dc:title>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>benthopelagic; freshwater</dc:description>
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