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	<dc:identifier>FB-8902</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8902</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Cichliformes</dwc:Order>
	<dwc:Family>Cichlidae</dwc:Family>
	<dwc:Genus>Heterotilapia</dwc:Genus>
	<dwc:ScientificName>Heterotilapia buettikoferi (Hubrecht, 1881)</dwc:ScientificName>
	<commonName xml:lang='da'>Hornet tilapia</commonName>
	<commonName xml:lang='de'>Zebratilapie</commonName>
	<commonName xml:lang='en'>Zebra tilapia</commonName>
	<commonName xml:lang='fi'>Seepratilapia</commonName>
	<commonName xml:lang=''>布氏罗非鱼</commonName>
	<commonName xml:lang=''>布氏非鯽</commonName>
	<commonName xml:lang=''>布氏非鲫</commonName>
	<synonym relationship='synonym'>Chromis buttikoferi Hubrecht, 1881</synonym>
	<synonym relationship='synonym'>Heterotilapia buttikoferi (Hubrecht, 1881)</synonym>
	<synonym relationship='synonym'>Tilapia ansorgii Boulenger, 1911</synonym>
	<synonym relationship='synonym'>Tilapia buttikoferi (Hubrecht, 1881)</synonym>
	<dcterms:created>1994-03-03</dcterms:created>
	<dcterms:modified>2024-03-22</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=2060&amp;speccode=8902'>Baensch, H.A. and R. Riehl0 Aquarien atlas. Band 2. Mergus, Verlag für Natur-und Heimtierkunde GmbH, Melle, Germany. 1216 p. (Ref. 2060)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4517&amp;speccode=8902'>Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5163&amp;speccode=8902'>Teugels, G.G. and D.F.E. Thys van den Audenaerde0 Tilapia. p. 482-508. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 5163)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7378&amp;speccode=8902'>Teugels, G.G. and D.F.E. Thys van den Audenaerde0 Cichlidae. p. 714-779. In C. Levêque, D. Paugy and G.G. Teugels (eds.) Faune des poissons d'eaux douces et saumâtres d'Afrique de l'Ouest. Tome 2. Coll. Faune Tropicale n° 28. Musée Royal de l'Afrique Centrale, Tervuren, Belgique and O.R.S.T.O.M., Paris, 902 p. (Ref. 7378)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=9932&amp;speccode=8902'>Pouyaud, L. and J.-F. Agnèse0 Phylogenetic relationships between 21 species of three tilapiine genera &lt;i&gt;Tilapia&lt;/i&gt;, &lt;i&gt;Sarotherodon&lt;/i&gt; and &lt;i&gt;Oreochromis&lt;/i&gt; using allozyme data. J. Fish Biol. 47(1):26-38. (Ref. 9932)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12818&amp;speccode=8902'>Anonymous0 Fish registrations within the museum database of the Vertebrate Section of the Royal Museum for Central Africa. MRAC, Tervuren, Belgium. (Ref. 12818)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=13331&amp;speccode=8902'>Paugy, D., K. Traoré and P.S. Diouf0 Faune ichtyologique des eaux douces d'Afrique de l'Ouest. p.35-66. In G.G. Teugels, J.F. Guégan and J.J. Albaret (eds.) Biological diversity of African fresh-and brackish water fishes. Geographical overviews presented at the PARADI Symposium, Senegal, 15-20 November 1993. Ann. Mus. R. Afr. Centr., Sci. Zool., 275:177 p. (Ref. 13331)</reference>
	<reference isbn='2-7099-I 340-2' url='http://www.fishbase.org/references/FBRefSummary.php?id=28587&amp;speccode=8902'>Diouf, P.S.0 Les peuplements de poissons des milieux estuariens de l'Afrique de l'Ouest: L'exemple de l'estuaire hyperhalin du Sine-Saloum. Université de Montpellier II. Thèses et Documents Microfiches No.156. ORSTOM, Paris. 267 p. (Ref. 28587)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31982&amp;speccode=8902'>Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=35508&amp;speccode=8902'>Anonymous0 The icthyological collection of the Zoological Museum Hamburg (ZMH). Division of Ichthyology and Herpetology, Zoological Museum Hamburg (ZMH). (Ref. 35508)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41414&amp;speccode=8902'>Anonymous0 Fish collection database of the American Museum of Natural History. American Museum of Natural History, Central Park West , NY 10024-5192, USA. (Ref. 41414)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51823&amp;speccode=8902'>Tan, B.C. and K.-S. Tan0 Singapore. p. 85-90. In N. Pallewatta, J.K. Reaser and A.T. Gutierrez (eds.) Invasive Alien Species in South-Southeast Asia: National Reports &amp; Directory of Resources. Global Invasive Species Programme, Cape Town, South Africa. 111p. (Ref. 51823)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52035&amp;speccode=8902'>Tang, B.Y.0 Fishes introduced to Singapore. Pers. com. Excel file attached to email dated 21/08/04. (Ref. 52035)</reference>
	<reference isbn='3-928819-33-X' url='http://www.fishbase.org/references/FBRefSummary.php?id=52307&amp;speccode=8902'>Lamboj, A.0 The cichlid fishes of Western Africa. Birgit Schmettkamp Verlag, Bornheim, Germany. 255 p. (Ref. 52307)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=53405&amp;speccode=8902'>Teugels, G.G. and D.F.E. Thys van den Audenaerde0 Cichlidae. p. 521-600. In D. Paugy, C. Lévêque and G.G Teugels (eds.) The fresh and brackish water fishes of West Africa Volume 2. Coll. faune et flore tropicales 40. Institut de recherche de développement, Paris, France, Muséum national d'histoire naturelle, Paris, France and Musée royal de l'Afrique Central, Tervuren, Belgium, 815p. (Ref. 53405)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57398&amp;speccode=8902'>Teugels, G.G. and D.F.E. Thys van den Audenaerde0 Cichlidae. p. 520-600. In C. Lévêque, D. Paugy and G.G. Teugels (eds.) Faune des poissons d'eaux douce et saumâtres de l'Afrique de l'Ouest, Tome 2. Coll. Faune et Flore tropicales 40. Musée Royal de l'Afrique Centrale, Tervuren, Belgique, Museum National d'Histoire Naturelle, Paris, France and Institut de Recherche pour le Développement, Paris, France. 815 p. (Ref. 57398)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=8902'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=74657&amp;speccode=8902'>Bartley, D.M. (comp./ed.)0 DIAS. Introduced species in fisheries and aquaculture: information for responsible use and control. CD-ROM. Rome (Italy): FAO. (Ref. 74657)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83882&amp;speccode=8902'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=93285&amp;speccode=8902'>Dunz, A.R. and U.K. Schliewen0 Molecular phylogeny and revised classification of the haplotilapiine cichlid fishes formerly referred to as "&lt;i&gt;Tilapia&lt;/i&gt;". Mol. Phylogenet. Evol. 68(1):64-80. (Ref. 93285)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=8902'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=126106&amp;speccode=8902'>Dean, E.M., A.R. Cooper, L. Wang, W. Daniel, S. David, C. Ernzen, K.B. Gido, E Hale, T.J. Haxton, W. Kelso, N. Leonard, C. Lido, J. Margraf, M. Porter, C. Pennock, D. Propst, J. Ross, M.D. Staudinger, G. Whelan and D.M. Infante0 The North American Freshwater Migratory Fish Database (NAFMFD): Characterizing the migratory life histories of freshwater fishes of Canada, the United States of Mexico. J. Biogeography 49:1193-1203. (Ref. 126106)</reference>
	<reference isbn='1849954984' url='http://www.fishbase.org/references/FBRefSummary.php?id=128868&amp;speccode=8902'>Beolens, B., M. Grayson and M. Watkins0 Eponym dictionary of Fishes. 	Dunbeath, Caithness (Scotland, UK): Whittles Publishing Limited, 1300 p. (Ref. 128868)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=129032&amp;speccode=8902'>Chan, J.C.F., A.H.F. Tsang, S.-M. Yau, T.C.H. Hui, A. Lau, H.H. Tan, B.W. Low, D. Dudgeon and J.H. Liew0 The non-native freshwater fishes of Hong Kong: diversity, distributions, and origins. Raffles Bull. Zool. 71:128-168. (Ref. 129032)</reference><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1994-03-03 00:00:00</dcterms:created>
	<dcterms:modified>2025-12-15 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>Adults occur only in larger rivers with rocky substrates, not found in smaller creeks; not specialized, but prefer a herbivorous diet; pair-bonding, biparental open substratum spawners holding very large territories; might spawn throughout the year (Ref. 52307).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=93285&amp;speccode=8902'>Dunz, A.R. and U.K. Schliewen 2013 Molecular phylogeny and revised classification of the haplotilapiine cichlid fishes formerly referred to as "&lt;i&gt;Tilapia&lt;/i&gt;". Mol. Phylogenet. Evol. 68(1):64-80. (Ref. 93285)</reference></dataObject><dataObject>
	<dc:identifier>FB-DiagnosticDescr-8902</dc:identifier>
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	<agent homepage='http://www.africamuseum.be#' logoURL='' role='recorder'>Tobias Musschoot</agent>
	<dcterms:created>2008-11-21 20:20:06</dcterms:created>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8902</dc:source>
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	<dc:description>Diagnosis: lower pharyngeal bone about as long as broad, and with anterior lamella shorter than toothed area; median pharyngeal teeth broadened when compared to the lateral teeth; 5-6 series of scales on cheeks; 4.5-6 scales between first dorsal fin spine and upper lateral line; dark vertical bars broader than lighter inter-spaces (Ref. 53405).Description: body (very) deep (Ref. 52307, 53405), its depth 44.4-52.2% of standard length (Ref. 53405). Frontal profile straight to concave (Ref. 52307). Outer jaw teeth bicuspid, outer jaw teeth and posterior pharyngeal teeth tricuspid; micro-gillrakers present; scales cycloid (Ref. 53405). Caudal fin slightly rounded (Ref. 52307).Coloration: ground colour yellowish-grey, with 7-8 broad, oblique (forward-directed) brownish-black bars, broader than light spaces between them and extending unto belly; 1st bar at level of eye, last on caudal peduncle; pectoral fins transparent, other fins brownish-black; pelvic fins with light anterior margin (Ref. 53405). Caudal fin with pale outer margin; dorsum of adult males sometimes pale bronze, females always somewhat lighter; lips pale blue; pelvic fins of adults always with elongated rays with a dark color and white anterior edge; older specimens tend to lose bars when comfortable, appearing totally dark gray to black, often with a bluish flush (Ref. 52307). Young: blackish area on chin; vertical bars on dorsal fin matching bars on sides of body which are less oblique than in adults; much darker bar at level of "tilapian" spot, reaching spinous part of anal fin, following bar (1st on caudal peduncle) extending unto soft parts of dorsal and anal fins; caudal fin with vertical bar at base and another at centre (Ref. 53405).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=53405&amp;speccode=8902'>Teugels, G.G. and D.F.E. Thys van den Audenaerde 2003 Cichlidae. p. 521-600. In D. Paugy, C. Lévêque and G.G Teugels (eds.) The fresh and brackish water fishes of West Africa Volume 2. Coll. faune et flore tropicales 40. Institut de recherche de développement, Paris, France, Muséum national d'histoire naturelle, Paris, France and Musée royal de l'Afrique Central, Tervuren, Belgium, 815p. (Ref. 53405)</reference></dataObject><dataObject>
	<dc:identifier>FB-Morphology-8902</dc:identifier>
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	<agent homepage='http://www.africamuseum.be#' logoURL='' role='recorder'>Tobias Musschoot</agent>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Dorsal spines (total): 13 - 15; Dorsal soft rays (total): 14 - 16;  Analspines: 3; Analsoft rays: 10 - 11</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=53405&amp;speccode=8902'>Teugels, G.G. and D.F.E. Thys van den Audenaerde 2003 Cichlidae. p. 521-600. In D. Paugy, C. Lévêque and G.G Teugels (eds.) The fresh and brackish water fishes of West Africa Volume 2. Coll. faune et flore tropicales 40. Institut de recherche de développement, Paris, France, Muséum national d'histoire naturelle, Paris, France and Musée royal de l'Afrique Central, Tervuren, Belgium, 815p. (Ref. 53405)</reference></dataObject><dataObject>
	<dc:identifier>FB-Distribution-8902</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1994-03-03 00:00:00</dcterms:created>
	<dcterms:modified>2025-12-15 00:00:00</dcterms:modified>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8902</dc:source>
	<dc:description>Africa:  lower reaches of coastal rivers from Guinea-Bissau (Geba and Corubal Rivers) to west Liberia (St. John River)(Ref. 53405).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5163&amp;speccode=8902'>Teugels, G.G. and D.F.E. Thys van den Audenaerde 1991 Tilapia. p. 482-508. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 5163)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-8902</dc:identifier>
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	<dc:title>Max. size</dc:title>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8902</dc:source>
	<dc:description>30.8 cm SL (male/unsexed; (Ref. 7378))</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7378&amp;speccode=8902'>Teugels, G.G. and D.F.E. Thys van den Audenaerde 1992 Cichlidae. p. 714-779. In C. Levêque, D. Paugy and G.G. Teugels (eds.) Faune des poissons d'eaux douces et saumâtres d'Afrique de l'Ouest. Tome 2. Coll. Faune Tropicale n° 28. Musée Royal de l'Afrique Centrale, Tervuren, Belgique and O.R.S.T.O.M., Paris, 902 p. (Ref. 7378)</reference></dataObject><dataObject>
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