FB-7348 http://www.fishbase.org/summary/SpeciesSummary.php?id=7348 Animalia Chordata Teleostei Perciformes/Serranoidei Epinephelidae Epinephelus Epinephelus howlandi (Günther, 1873) Blacksaddle grouper Blacksaddle rockcod Blacksaddled cod Howland island rock cod kawakawa Mérou sellé noir Lokos Hiregurohata Kabro Mero montura negra Kusele Nefu Tumuvutuvo 斑吻石斑魚 斑吻石斑鱼 荷氏石斑魚 荷氏石斑鱼 花杉頭 Epinephelus corallicola (non Valenciennes, 1828) Epinephelus macrospilos (non Bleeker, 1855) Epinephelus spilotus Schultz, 1953 Serranus howlandi Günther, 1873 1993-02-12 2015-04-23 Coppola, S.R., W. Fischer, L. Garibaldi, N. Scialabba and K.E. Carpenter0 SPECIESDAB: Global species database for fishery purposes. User's manual. FAO Computerized Information Series (Fisheries). No. 9. Rome, FAO. 103 p. (Ref. 171) Hiatt, R.W. and D.W. Strasburg0 Ecological relationships of the fish fauna on coral reefs of the Marshall Islands. Ecol. Monogr. 30(1):65-127. (Ref. 275) Schultz, L.P., E.S. Herald, E.A. Lachner, A.D. Welander and L.P. Woods0 Fishes of the Marshall and Marianas Islands. Vol. I. Families from Asymmetrontidae through Siganidae. Bull. U.S. Natl. Mus. 202(1):685. (Ref. 1717) Eschmeyer, W.N.0 Catalog of the genera of recent fishes. California Academy of Sciences, San Francisco, USA. 697 p. (Ref. 1830) Randall, J.E., G.R. Allen and R.C. Steene0 Fishes of the Great Barrier Reef and Coral Sea. University of Hawaii Press, Honolulu, Hawaii. 506 p. (Ref. 2334) Myers, R.F.0 Micronesian reef fishes: A practical guide to the identification of the inshore marine fishes of the tropical central and western Pacific. First Edition. Coral Graphics, Barrigada, Guam. 298 p. (Ref. 4538) Randall, J.E. and P.C. Heemstra0 Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with descriptions of five new species. Indo-Pac. Fish. (20):332 p. (Ref. 4787) Heemstra, P.C. and J.E. Randall0 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222) Anonymous0 Computerized catalog of the fish collection. California Academy of Sciences, San Francisco, California. (Ref. 5515) Kailola, P.J.0 The fishes of Papua New Guinea: a revised and annotated checklist. Vol. II Scorpaenidae to Callionymidae. Research Bulletin No. 41, Research Section, Dept. of Fisheries and Marine Resources, Papua New Guinea. (Ref. 6192) Katayama, M.0 Family Serranidae. p. 126-138. In H. Masuda, K. Amaoka, C. Araga, T. Uyeno, and T. Yoshino (eds). The fishes of the Japanese archipelago. Second Edition. Tokai University Press, Tokyo. 456 p. (Ref. 6854) Katayama, M.0 Fauna Japonica. Serranidae (Pisces). Tokyo, Tokyo News Service. 189 p., 86 pls. (Ref. 6864) Francis, M.P.0 Checklist of the coastal fishes of Lord Howe, Norfolk, and Kermadec Islands, Southwest Pacific Ocean. Pac. Sci. 47(2):136-170. (Ref. 8879) Masuda, H. and G.R. Allen0 Meeresfische der Welt - Groß-Indopazifische Region. Tetra Verlag, Herrenteich, Melle. 528 p. (Ref. 9137) Lieske, E. and R. Myers0 Collins Pocket Guide. Coral reef fishes. Indo-Pacific & Caribbean including the Red Sea. Haper Collins Publishers, 400 p. (Ref. 9710) Joannot, P.0 List of fishes in the Noumea aquarium. Unpublished. (Ref. 13603) Foale, S.0 What's in a name? An analysis of the West Nggela (Solomon Islands) fish taxonomy. SPC Traditional Marine Resource Management and Knowledge Information Bulletin No. 9:3-19. (Ref. 26367) Letourneur, Y., M. Kulbicki and P. Labrosse0 Length-weight relationships of fish from coral reefs and lagoons of New Caledonia, southwestern Pacific Ocean: an update. Naga, ICLARM Q. 21(4):39-46. (Ref. 26587) Lee, C. and Y. Sadovy0 A taste for live fish: Hong Kong's live reef fish market. Naga, ICLARM Q. 21(2):38-42. (Ref. 27253) Randall, J.E.0 Randall's tank photos. 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Bishop Museum, 1525 Bernice Street, Honolulu, Hawai`I, 96817-0916 USA. (Ref. 31667) Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982) Lau, P.P.F. and L.W.H. Li0 Identification guide to fishes in the live seafood trade of the Asia-Pacific region. World Wide Fund for Nature, Hong Kong. 137 p. (Ref. 34166) Anonymous0 The icthyological collection of the Zoological Museum Hamburg (ZMH). Division of Ichthyology and Herpetology, Zoological Museum Hamburg (ZMH). (Ref. 35508) Randall, J.E. and K.K.P. Lim (eds.)0 A checklist of the fishes of the South China Sea. Raffles Bull. Zool. Suppl. (8):569-667. (Ref. 36648) Myers, R.F.0 Micronesian reef fishes: a comprehensive guide to the coral reef fishes of Micronesia, 3rd revised and expanded edition. Coral Graphics, Barrigada, Guam. 330 p. (Ref. 37816) Anonymous0 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes. (Ref. 38732) Heemstra, P.C. and J.E. Randall0 Serranidae. p. 2442-2547. In K.E. Carpenter and V.H. Niem (eds.) FAO species identification guide for fishery purposes. The living marine resources of the Western Central Pacific. Vol. 4. Bony fishes part 2 (Mugilidae to Carangidae). Rome, FAO. 2069-2790 p. (Ref. 39231) Anonymous0 Fish collection database of the American Museum of Natural History. American Museum of Natural History, Central Park West , NY 10024-5192, USA. (Ref. 41414) Allen, G.R.0 Reef fishes of Milne Bay Province, Papua New Guinea. In T. Werner and G. Allen (eds). A rapid biodiversity assessment of the coral reefs of Milne Bay Province, Papua New Guinea. RAP Working Papers 11, Conservation International, Washington, D.C. (Ref. 41464) Anonymous0 Fish collection of the Royal Ontario Museum. Royal Ontario Museum. (Ref. 47438) Hardy, J.D. Jr. (comp.)0 Coral reef fish species. NOAANational Oceanographic Data Center. NODC Coral Reef Data and Information Management System. USA. 537 p. (Ref. 48497) Kuiter, R.H. and T. Tonozuka0 Pictorial guide to Indonesian reef fishes. Part 1. Eels- Snappers, Muraenidae - Lutjanidae. Zoonetics, Australia. 1-302. (Ref. 48635) Randall, J.E.0 Reef and shore fishes of the South Pacific. New Caledonia to Tahiti and the Pitcairn Islands. University of Hawaii Press, Honolulu, Hawaii. 720 p. (Ref. 54980) Kulbicki, M., N. Guillemot and M. Amand0 A general approach to length-weight relationships for New Caledonian lagoon fishes. Cybium 29(3):235-252. (Ref. 55787) Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108) Hoese, D.F., D.J. Bray, J.R. Paxton and G.R. Allen0 Fishes. In Beasley, O.L. and A. Wells (eds.) Zoological Catalogue of Australia. Volume 35 Australia: ABRS & CSIRO Publishing, 2178 p. (Ref. 75154) Thaman, R.R., T. Fong and A. Balawa0 Ilava Ni Navakavu: Finfishes of Vanua Navakavu, Viti Levu, Fiji Islands. SPRH-FIO Biodiversity and Ethnobiodiversity Report No. 4, The University of the South Pacific, Suva, Fiji Islands. (Ref. 79890) Kronen, M., E. Tardy, P. Boblin, L. Chapman, F. Lasi, K. Pakoa, L. Vigliola, K. Friedman, F. Magron and S. Pinca0 Wallis and Futuna Country Report: Profiles and results from survey work at Vailala, Halalo, Leava and Vele. Secretariat of the Pacific Community. Pacific Regional Oceanic and Coastal Fisheries Development Programme (PROCFish/C/CoFish). Aug-Dec 2005 and Mar 2006. (Ref. 84356) Seeto, J. and W.J. Baldwin0 A checklist of the fishes of Fiji and a bibliography of Fijian fish. Division of Marine Studies Technical Report 1/2010. The University of the South Pacific. Suva, Fiji. 102 p. (Ref. 85272) Fricke, R., M. Kulbicki and L. Wantiez0 Checklist of the fishes of New Caledonia, and their distribution in the Southwest Pacific Ocean (Pisces). Stuttgarter Beiträge zur Naturkunde A, Neue Serie 4:341-463. (Ref. 86942) Craig, M.T., YJ. Sadovy de Mitcheson and P.C. Heemstra0 Groupers of the world: a field and market guide. North America: CRC Press/Taylor and Francis Group, xix, 356 p., A47 pages appendix. (Ref. 89707) Donaldson, T.J.0 Fishes of the remote Southwest Palau Islands: a zoogeographic perspective. Pac. Sci. 50(3):285-308. (Ref. 99878) Kerr, V.C.0 A rapid marine biodiversity assessment of the coral reefs of the northwest lagoon, between Koumac and Yandé, Province Nord, New Caledonia. BioOne Research Evolved:160-178. (Ref. 101627) Kanikawa, K.T., E. Cruz, T.E. Essington, J. Hospital, J.K.T. Brodziak and T.A. Branch0 Length-weight relationships for 85 fish species from Guam. J. Appl. Ichthyol. 31:1171-1174. (Ref. 107958) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from htttps://www.fao.org/fishery/collection/asfis/en [accessed May, 2018] (Ref. 118272) Matthews, T., J. Gourley, A. Flores, M. Ramon and M. Trianni0 Length-weight relationships for 83 reef and bottomfish species from the commonwealth of the Northern Mariana Islands. Pacific Islands Fish. Sci. Cent. Admin. Rep. H-19-04, 7 p. (Ref. 125598) Price, S.A., S.T. Friedman, K.A. Corn, O. Larouche, K. Brockelsby, A.J. Lee, M. Nagaraj, N.G. Bertrand, M. Danao, M.C. Coyne, J.R. Estrada, R. Friedman, E. Hoeft, M. Iwan, D. Gross, J.H. Kao, B. Landry, M.J. Linares, C. McGlinn and …0 FishShapes v1: Functionally relevant measurements of teleost shape and size on three dimensions. Ecology 103(e3829):1-2. (Ref. 137577) FB-pic-7348-2259 http://purl.org/dc/dcmitype/StillImage image/jpeg John E. Randall 1995-01-06 2010-07-13 http://creativecommons.org/licenses/by-nc/3.0/ This work is licensed under a Creative Commons Attribution-Noncommercial 3.0 Unported License. Image can be freely copied and altered but may be used only for non-commercial purposes. Original author and source must be properly acknowledged WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=7348 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Morphology http://www.fishbase.us/images/species/Ephow_u0.jpg http://www.fishbase.us/images/thumbnails/jpg/tn_Ephow_u0.jpg FB-pic-7348-9549 http://purl.org/dc/dcmitype/StillImage image/jpeg John E. Randall 1996-03-03 2000-11-08 http://creativecommons.org/licenses/by-nc/3.0/ This work is licensed under a Creative Commons Attribution-Noncommercial 3.0 Unported License. Image can be freely copied and altered but may be used only for non-commercial purposes. Original author and source must be properly acknowledged WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=7348 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution http://www.fishbase.us/images/species/Ephow_u1.jpg http://www.fishbase.us/images/thumbnails/jpg/tn_Ephow_u1.jpg FB-pic-7348-9550 http://purl.org/dc/dcmitype/StillImage image/jpeg John E. Randall 1996-03-03 2000-11-08 http://creativecommons.org/licenses/by-nc/3.0/ This work is licensed under a Creative Commons Attribution-Noncommercial 3.0 Unported License. Image can be freely copied and altered but may be used only for non-commercial purposes. Original author and source must be properly acknowledged WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=7348 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Uses market Locality: Naha, Okinawa, Ryukyu Is.. http://www.fishbase.us/images/species/Ephow_u2.jpg http://www.fishbase.us/images/thumbnails/jpg/tn_Ephow_u2.jpg Locality: Naha, Okinawa, Ryukyu Is.. FB-pic-7348-9551 http://purl.org/dc/dcmitype/StillImage image/jpeg John E. Randall 1996-03-03 2000-11-08 http://creativecommons.org/licenses/by-nc/3.0/ This work is licensed under a Creative Commons Attribution-Noncommercial 3.0 Unported License. Image can be freely copied and altered but may be used only for non-commercial purposes. Original author and source must be properly acknowledged WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=7348 http://rs.tdwg.org/ontology/voc/SPMInfoItems#TrophicStrategy http://www.fishbase.us/images/species/Ephow_u3.jpg http://www.fishbase.us/images/thumbnails/jpg/tn_Ephow_u3.jpg FB-pic-7348-11205 http://purl.org/dc/dcmitype/StillImage image/gif FAO 1996-05-25 1996-05-25 http://creativecommons.org/licenses/by-nc/3.0/ This work is licensed under a Creative Commons Attribution-Noncommercial 3.0 Unported License. Image can be freely copied and altered but may be used only for non-commercial purposes. Original author and source must be properly acknowledged WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=7348 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size http://www.fishbase.us/images/species/Ephow_u0.gif http://www.fishbase.us/images/thumbnails/gif/tn_Ephow_u0.gif FB-GeneralDescr-7348 http://purl.org/dc/dcmitype/Text text/html Estelita Emily Capuli 1993-02-12 00:00:00 2015-04-23 00:00:00 Biology http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=7348 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats Usually found over rocky areas or on coral reefs. Feeds on fishes and crustaceans (shrimps and crab) with occasional molluscs or worms (Ref. 89707). Nothing has been published on its biology. Appears to be very rare and therefore of little commercial importance. In Hong Kong live fish markets (Ref. 27253). Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222) FB-DiagnosticDescr-7348 http://purl.org/dc/dcmitype/Text text/html Rodolfo B. Reyes 1994-06-13 00:00:00 2013-10-11 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=7348 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat Distinguished by the following characteristics: head and body pale grey or brownish grey, covered with small brownish black spots with diffuse edges, separated from adjacent spots by spaces equal to or greater than width of the spots; body with black saddle blotch , smaller black saddle blotch on the caudal peduncle; dusky margins of the soft dorsal, caudal and anal fins, with white line along edge; median and pelvic fins with dark spots; dusky pectoral fins with few dark spots basally; body depth contained 2.9-3.3 times in SL; HL 2.2-2.5 times in SL; flat interorbital area, convex dorsal head profile; rounded preopercle, slightly enlarged serrae at angle; maxilla reaches past vertical at rear edge of eye; 2-4 rows of teeth on the midlateral part of lower jaw; pyloric caeca 13 (Ref. 89707). Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222) FB-Morphology-7348 http://purl.org/dc/dcmitype/Text text/html Rodolfo B. Reyes 1994-06-13 00:00:00 2013-10-11 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ http://www.fishbase.org/summary/SpeciesSummary.php?id=7348 Dorsal spines (total): 11; Dorsal soft rays (total): 15 - 17; Analspines: 3; Analsoft rays: 8 Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222) FB-Distribution-7348 http://purl.org/dc/dcmitype/Text text/html Estelita Emily Capuli 1993-02-12 00:00:00 2018-01-18 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=7348 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Morphology Western Pacific: Ryukyu Islands south to Lord Howe Island and east to Howland Island and Samoa. Often misidentified as Epinephelus macrospilos or Epinephelus corallicola. Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222) FB-Uses-7348 http://purl.org/dc/dcmitype/Text text/html Estelita Emily Capuli 1993-02-12 00:00:00 2015-04-23 00:00:00 Importance http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=7348 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution fisheries: subsistence fisheries; aquarium: public aquariums Coppola, S.R., W. Fischer, L. Garibaldi, N. Scialabba and K.E. Carpenter 1994 SPECIESDAB: Global species database for fishery purposes. User's manual. FAO Computerized Information Series (Fisheries). No. 9. Rome, FAO. 103 p. (Ref. 171) Joannot, P. 1997 List of fishes in the Noumea aquarium. Unpublished. (Ref. 13603) FB-TrophicStrategy-7348 http://purl.org/dc/dcmitype/Text text/html Pascualita Sa-a 1997-03-25 00:00:00 2008-11-19 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ howlandi Found inshore (Ref. 75154). Feeds on fish and benthic invertebrates (Ref. 275). Hiatt, R.W. and D.W. Strasburg 1960 Ecological relationships of the fish fauna on coral reefs of the Marshall Islands. Ecol. Monogr. 30(1):65-127. (Ref. 275) FB-Size-7348 http://purl.org/dc/dcmitype/Text text/html Estelita Emily Capuli 1993-02-12 00:00:00 2015-04-23 00:00:00 Max. size http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=7348 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Migration 55.0 cm TL (male/unsexed; (Ref. 48635)) Kuiter, R.H. and T. Tonozuka 2001 Pictorial guide to Indonesian reef fishes. Part 1. Eels- Snappers, Muraenidae - Lutjanidae. Zoonetics, Australia. 1-302. (Ref. 48635) FB-Threats-2 http://purl.org/dc/dcmitype/Text text/html Belen Acosta 1990-10-18 00:00:00 2024-03-07 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=2 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006. FB-Habitat-7348 http://purl.org/dc/dcmitype/Text text/html Estelita Emily Capuli 1993-02-12 00:00:00 2015-04-23 00:00:00 Environment http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=7348 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats reef-associated; marine; depth range 1 - 37 m (Ref. 86942) Fricke, R., M. Kulbicki and L. Wantiez 2011 Checklist of the fishes of New Caledonia, and their distribution in the Southwest Pacific Ocean (Pisces). Stuttgarter Beiträge zur Naturkunde A, Neue Serie 4:341-463. (Ref. 86942)