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	<dc:identifier>FB-15484</dc:identifier>
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	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Elasmobranchii</dwc:Class>
	<dwc:Order>Myliobatiformes</dwc:Order>
	<dwc:Family>Dasyatidae</dwc:Family>
	<dwc:Genus>Brevitrygon</dwc:Genus>
	<dwc:ScientificName>Brevitrygon walga (Müller &amp; Henle, 1841)</dwc:ScientificName>
	<commonName xml:lang='cs'>Trnucha Toshova</commonName>
	<commonName xml:lang='da'>Sortplettet piskerokke</commonName>
	<commonName xml:lang='en'>Black-edged whipray</commonName>
	<commonName xml:lang='en'>Black-spotted stingray</commonName>
	<commonName xml:lang='en'>Black-spotted whip ray</commonName>
	<commonName xml:lang='en'>Blackspotted whipray</commonName>
	<commonName xml:lang='en'>Black-spotted whipray</commonName>
	<commonName xml:lang='en'>Coachwhip ray</commonName>
	<commonName xml:lang='en'>Toshs longtail ray</commonName>
	<commonName xml:lang='en'>Wulura</commonName>
	<commonName xml:lang='et'>Toshi nuutrai</commonName>
	<commonName xml:lang='jv'>Pari tutul</commonName>
	<commonName xml:lang='ms'>Pari bintik hitam</commonName>
	<commonName xml:lang='ms'>Pari rimau</commonName>
	<commonName xml:lang='th'>Krabane Lai-mang-won</commonName>
	<commonName xml:lang=''>Pari aer</commonName>
	<commonName xml:lang=''>Zwartgevlekte pijlstaartrog</commonName>
	<commonName xml:lang=''>斑点窄尾魟</commonName>
	<commonName xml:lang=''>斑點窄尾魟</commonName>
	<synonym relationship='synonym'>Himantura toshi Whitley, 1939</synonym>
	<synonym relationship='misapplied name'>Himantura uarnak (non Forsskål, 1775)</synonym>
	<dcterms:created>1995-10-25</dcterms:created>
	<dcterms:modified>2024-05-23</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=15487'>Breder, C.M. and D.E. Rosen0 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference>
	<reference isbn='0 643 05143 0' url='http://www.fishbase.org/references/FBRefSummary.php?id=6871&amp;speccode=15487'>Last, P.R. and J.D. Stevens0 Sharks and rays of Australia. CSIRO, Australia. 513 p. (Ref. 6871)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=6932&amp;speccode=15487'>Salini, J.P., S.J.M. Blaber and D.T`. Brewer0 Diets of trawled predatory fish of the Gulf of Carpentaria, Australia, with particular reference to predation on prawns. Aust. J. Mar. Freshwat. Res. 45(3):397-411. (Ref. 6932)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=9840&amp;speccode=15487'>Last, P.R. and L.J.V. Compagno0 Dasyatididae. Stingrays. p. 1479-1505. In K.E. Carpenter and V.H. Niem (eds.) FAO species identification guide for fishery purposes. The living marine resources of the Western Central Pacific. Vol. 3. Batoid fishes, chimaeras and bony fishes part 1 (Elopidae to Linophrynidae). FAO, Rome. (Ref. 9840)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=15487'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='0-8018-6048-2' url='http://www.fishbase.org/references/FBRefSummary.php?id=35766&amp;speccode=15487'>Compagno, L.J.V.0 Checklist of living elasmobranchs. p. 471-498. In W.C. Hamlett (ed.) Sharks, skates, and rays: the biology of elasmobranch fishes. Johns Hopkins University Press, Maryland. (Ref. 35766)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=38475&amp;speccode=15487'>Last, P.R. and L.J.V. Compagno0 Dasyatididae. Stingrays. p. 1479-1505. In K.E. Carpenter and V.H. Niem (eds.) FAO species identification guide for fishery purposes. The living marine resources of the Western Central Pacific. Vol. 3. Batoid fishes, chimaeras and bony fishes part 1 (Elopidae to Linophrynidae). FAO, Rome. (Ref. 38475)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41414&amp;speccode=15487'>Anonymous0 Fish collection database of the American Museum of Natural History. American Museum of Natural History, Central Park West , NY 10024-5192, USA. (Ref. 41414)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=43454&amp;speccode=15487'>Bor, P.H.F.0 Nederlandse naamlijst van de recente haaien en roggen (Chondrichthyes: Elasmobranchii) van de wereld. World Wide Web electronic publication www.rajidae.tmfweb.nl, version (05/2002). (Ref. 43454)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=50449&amp;speccode=15487'>Dulvy, N.K. and J.D. Reynolds0 Evolutionary transitions among egg-laying, live-bearing and maternal inputs in sharks and rays. Proc. R. Soc. Lond., Ser. B: Biol. Sci. 264:1309-1315. (Ref. 50449)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=50831&amp;speccode=15487'>Seafood Services Australia0 Australian fish names list. http://www.seafoodservices.com.au/, version 16 September 2003. (Ref. 50831)</reference>
	<reference isbn='80-7036-139-5' url='http://www.fishbase.org/references/FBRefSummary.php?id=51055&amp;speccode=15487'>Hanel, L. and J. Novák0 České názvy zivočichů V. Ryby a ryboviti obratlovci (Pisces) 3., maloústí (Gonorhynchiformes) - máloostní (Cypriniformes). Národní muzeum (zoologické oddělení), Praha. (Ref. 51055)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51471&amp;speccode=15487'>Carl, H.0 Danish fish names. Zoological Museum of Copenhagen. Unpublished. (Ref. 51471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=53392&amp;speccode=15487'>Ahmad, A., A.A. Abdul Haris Hilmi, A.C. Gambang, S. Ahemad and A.R. Solahuddin (eds.)0 Elasmobranch resources, utilization, trade and management in Malaysia. Marine Fishery Resources Development and Management Department, Southeast Asian Fisheries Development Center. (Ref. 53392)</reference>
	<reference isbn='1 86320 516 0' url='http://www.fishbase.org/references/FBRefSummary.php?id=58048&amp;speccode=15487'>White, W.T., P.R. Last, J.D. Stevens, G.K. Yearsley, Fahmi and Dharmadi0 Economically important sharks and rays of Indonesia. [Hiu dan pari yang bernilai ekonomis penting di Indonesia]. Australian Centre for International Agricultural Research, Canberra, Australia. (Ref. 58048)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=15487'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=75154&amp;speccode=15487'>Hoese, D.F., D.J. Bray, J.R. Paxton and G.R. Allen0 Fishes. In Beasley, O.L. and A. Wells (eds.) Zoological Catalogue of Australia. Volume 35 Australia: ABRS &amp; CSIRO Publishing, 2178 p. (Ref. 75154)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=75549&amp;speccode=15487'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis, 29 April 2008. (Ref. 75549)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83882&amp;speccode=15487'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882)</reference>
	<reference isbn='978-983-9114-46-1' url='http://www.fishbase.org/references/FBRefSummary.php?id=85449&amp;speccode=15487'>Department of Fisheries Malaysia0 Valid local name of Malaysian marine fishes. Department of Fisheries Malaysia. Ministry of Agriculture and Agro-based Industry. 180 p. (Ref. 85449)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=15487'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='9780987260000' url='http://www.fishbase.org/references/FBRefSummary.php?id=90102&amp;speccode=15487'>Allen, G.R. and M.V. Erdmann0 Reef fishes of the East Indies. Perth, Australia: Universitiy of Hawai'i Press, Volumes I-III. Tropical Reef Research. (Ref. 90102)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=15487'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=15487'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105471&amp;speccode=15487'>eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=114953&amp;speccode=15487'>Last, P.R., W.T. White, M.R. de Carvalho, B. Séret, M.F.W. Stehmann and G.J.P. Naylor0 Rays of the world. CSIRO Publishing, Comstock Publishing Associates. i-ix + 1-790. (Ref. 114953)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115257&amp;speccode=15487'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257)</reference>
	<reference isbn='1849954984' url='http://www.fishbase.org/references/FBRefSummary.php?id=128868&amp;speccode=15487'>Beolens, B., M. Grayson and M. Watkins0 Eponym dictionary of Fishes. 	Dunbeath, Caithness (Scotland, UK): Whittles Publishing Limited, 1300 p. (Ref. 128868)</reference><dataObject>
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	<dcterms:created>2001-05-02</dcterms:created>
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	<mediaURL>http://www.fishbase.us/images/species/Hitos_u0.gif</mediaURL>
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	<agent homepage='' logoURL='' role='recorder'>Cristina V. Garilao</agent>
	<dcterms:created>1995-10-25 00:00:00</dcterms:created>
	<dcterms:modified>2024-05-23 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=15487</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Demersal on soft bottoms on the continental shelf at depths of 10-140m (Ref.58048). Juveniles (20-22 cm WD) are free swimming.  Feeds on prawns, small fishes and crustaceans (Ref. 9840).  Ovoviviparous (Ref. 50449). Common by-catch of bottom trawl and beach seine fisheries in the Arafura Sea. Utilized for its meat, skin (high value) and cartilage (Ref.58048). </dc:description>
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	<reference isbn='0 643 05143 0' url='http://www.fishbase.org/references/FBRefSummary.php?id=6871&amp;speccode=15487'>Last, P.R. and J.D. Stevens 1994 Sharks and rays of Australia. CSIRO, Australia. 513 p. (Ref. 6871)</reference></dataObject><dataObject>
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	<dcterms:created>1995-10-25 00:00:00</dcterms:created>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>Western Pacific:  Australia and Papua New Guinea.  This species has been misidentified by most ichthyologists as a color form of Himantura uarnak (Ref. 6871). Also Ref. 114953.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=114953&amp;speccode=15487'>Last, P.R., W.T. White, M.R. de Carvalho, B. Séret, M.F.W. Stehmann and G.J.P. Naylor 2016 Rays of the world. CSIRO Publishing, Comstock Publishing Associates. i-ix + 1-790. (Ref. 114953)</reference></dataObject><dataObject>
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	<dcterms:created>1995-10-25 00:00:00</dcterms:created>
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	<dc:title>Importance</dc:title>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>fisheries: minor commercial; price category: low; price reliability: very questionable: based on ex-vessel price for species in this family</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=9840&amp;speccode=15487'>Last, P.R. and L.J.V. Compagno 1999 Dasyatididae. Stingrays. p. 1479-1505. In K.E. Carpenter and V.H. Niem (eds.) FAO species identification guide for fishery purposes. The living marine resources of the Western Central Pacific. Vol. 3. Batoid fishes, chimaeras and bony fishes part 1 (Elopidae to Linophrynidae). FAO, Rome. (Ref. 9840)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1995-10-25 00:00:00</dcterms:created>
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	<dcterms:rightsHolder> FAO</dcterms:rightsHolder>
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	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=15487&amp;StockCode=14735</dc:source>
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	<dc:description>Exhibit ovoviparity (aplacental viviparity), with embryos feeding initially on yolk, then receiving additional nourishment from the mother by indirect absorption of uterine fluid enriched with mucus, fat or protein through specialised structures (Ref. 50449). Distinct pairing with embrace (Ref. 205).  Born at 20-22 cm WD (Ref.58048).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=15487'>Breder, C.M. and D.E. Rosen 1966 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference></dataObject><dataObject>
	<dc:identifier>FB-TrophicStrategy-15487</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1999-07-21 00:00:00</dcterms:created>
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	<dcterms:rightsHolder>John E. Randall</dcterms:rightsHolder>
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	<dc:source>toshi</dc:source>
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	<dc:description>Found on the continental shelf (Ref. 9840, 75154).  Juveniles (20-22 cm WD) are free swimming.  Feed on prawns, small fishes and crustaceans (Ref. 9840).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=6932&amp;speccode=15487'>Salini, J.P., S.J.M. Blaber and D.T`. Brewer 1994 Diets of trawled predatory fish of the Gulf of Carpentaria, Australia, with particular reference to predation on prawns. Aust. J. Mar. Freshwat. Res. 45(3):397-411. (Ref. 6932)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-15487</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Cristina V. Garilao</agent>
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	<dc:title>Max. size</dc:title>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=15487</dc:source>
	<subject></subject>
	<dc:description>86.0 cm WD (male/unsexed; (Ref. 58048))</dc:description>
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	<reference isbn='1 86320 516 0' url='http://www.fishbase.org/references/FBRefSummary.php?id=58048&amp;speccode=15487'>White, W.T., P.R. Last, J.D. Stevens, G.K. Yearsley, Fahmi and Dharmadi 2006 Economically important sharks and rays of Indonesia. [Hiu dan pari yang bernilai ekonomis penting di Indonesia]. Australian Centre for International Agricultural Research, Canberra, Australia. (Ref. 58048)</reference></dataObject><dataObject>
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	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
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	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-15487</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Cristina V. Garilao</agent>
	<dcterms:created>1995-10-25 00:00:00</dcterms:created>
	<dcterms:modified>2024-05-23 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>John E. Randall</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=15487</dc:source>
	<subject></subject>
	<dc:description>demersal; marine; depth range 10 - 140 m (Ref. 6871)</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='0 643 05143 0' url='http://www.fishbase.org/references/FBRefSummary.php?id=6871&amp;speccode=15487'>Last, P.R. and J.D. Stevens 1994 Sharks and rays of Australia. CSIRO, Australia. 513 p. (Ref. 6871)</reference></dataObject>
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