Actinopteri (ray-finned fishes) > Mulliformes
(Goatfishes) > Mullidae
Etymology: Upeneus: Greek, ypene, -es = upper lip (Ref. 45335); nigromarginatus: Species name refers to the black margins along both dorsal fins and the dorsal lobe of the caudal fin.
Environment: milieu / climate zone / depth range / distribution range
Marine; demersal; depth range 40 - 80 m (Ref. 97569). Tropical; 20°N - 5°N, 117°E - 127°E
Western Pacific: Philippines.
Size / Weight / Age
Maturity: Lm ?  range ? - ? cm
Max length : 27.0 cm SL male/unsexed; (Ref. 108462)
Morphology | Morphometrics
soft rays: 6. This species is distinguished by the following characters: D VIII,9, with first dorsal spine small, second dorsal spine longest, last dorsal soft ray branched to base; A I,6 (3 paratypes with unusual two spines and first spine short and thin, second spine up to two-third of first soft ray, first anal soft ray segmented and unbranched, last ray branched to base; pectoral fin rays 16, upper 2 rays unbranched; pelvic fin I,5; principal caudal fin rays 15 (15-16), with uppermost and lowermost rays unbranched; lateral line scales 36 (36-37), five scale rows between dorsal fins; total gill rakers on first gill arch 25 (25-26), with gill rakers on upper and lower limb of first arch 8 + 17 (7-9 + 17-18), and several anterior rakers undeveloped but visible on both limbs (Ref. 97569).
Life cycle and mating behavior
Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae
Bos, A.R., 2014. Upeneus nigromarginatus, a new species of goatfish (Perciformes: Mullidae) from the Philippines. Raffles Bulletin of Zoology 62: 745-753. (Ref. 97569)
IUCN Red List Status (Ref. 123251)
CITES (Ref. 123416)
Threat to humans
Common namesSynonymsMetabolismPredatorsEcotoxicologyReproductionMaturitySpawningSpawning aggregationFecundityEggsEgg development
ReferencesAquacultureAquaculture profileStrainsGeneticsAllele frequenciesHeritabilityDiseasesProcessingNutrientsMass conversion
Estimates based on models
Phylogenetic diversity index (Ref. 82805
= 0.5000 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.01380 (0.00803 - 0.02372), b=3.18 (3.04 - 3.32), in cm total length, based on LWR estimates for this species & Genus-body shape (Ref. 93245
Trophic level (Ref. 69278
): 3.6 ±0.5 se; based on size and trophs of closest relatives
Resilience (Ref. 120179
): Medium, minimum population doubling time 1.4 - 4.4 years (Preliminary K or Fecundity.).
Vulnerability (Ref. 59153
): Low to moderate vulnerability (34 of 100) .