Environment: milieu / climate zone / depth range / distribution range
Ecology
Freshwater; demersal. Temperate; 46°N - 35°N
North America: Pacific Slope drainages from lower Columbia River drainage in Washington to Morro Bay in California (including Sacramento-San Joaquin River drainage except upper Pit River), USA; absent in Rogue and Klamath River drainages in southern Oregon and northern California.
Size / Weight / Age
Maturity: Lm ? range ? - ? cm
Max length : 11.0 cm TL male/unsexed; (Ref. 5723); common length : 6.0 cm TL male/unsexed; (Ref. 12193)
Inhabits sand and gravel riffles of headwaters and creeks; also in sand-gravel runs and backwaters of small to large rivers (Ref. 5723).
Life cycle and mating behavior
Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae
Page, L.M. and B.M. Burr, 1991. A field guide to freshwater fishes of North America north of Mexico. Houghton Mifflin Company, Boston. 432 p. (Ref. 5723)
IUCN Red List Status (Ref. 130435: Version 2025-2)
Threat to humans
Harmless
Human uses
Fisheries: of no interest
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