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Gobiiformes (Gobies) > 
Gobiidae (Gobies) > Gobionellinae						
							
							Etymology: Gnatholepis: Greek, gnathos = jaw + Greek,lepis = scale (Ref. 45335).
More on author: Bleeker.						
					
				
					Environment: milieu / climate zone / Mức độ sâu / distribution range					
						Sinh thái học					
				
				
				
					Biển; Thuộc về nước lợ Cùng sống ở rạn san hô;  Mức độ sâu 1 - 46 m (Ref. 11441). Tropical				
				 
			
			
			
				
				
				
					Indo-Pacific:  Persian Gulf (Ref.80050) and Red Sea and East Africa to the Hawaiian, Marquesan and Society islands.
				
				
			
			
				
					Bộ gần gũi / Khối lượng (Trọng lượng) / Age
				
				
				
					Chín muồi sinh dục: Lm ?  range ? - ? cm
 Max length : 8.4 cm SL con đực/không giới tính; (Ref. 41653)				
				 
			
			
						
				
				
				
					Các tia vây lưng cứng (tổng cộng) : 6 - 7; Các vây lưng mềm (tổng cộng) : 11; Tia cứng vây hậu môn: 1; Tia mềm vây hậu môn: 11. Distinguished by the following characteristics: ctenoid scales on head, predorsal midline and pectoral fin base, distinct flap present at end of lower lip; at least three dark spots along first spine of first and second dorsal fins, followed by at least three rows of dark spots or short streaks, pectoral fin with rows of fine dark and white speckles; second dorsal and anal fin rays nearly always I,11; pectoral rays 14-17, modally 16; lateral scales 25-29, modally 27; predorsal scales 7-13, usually 9-10, mostly ctenoid, may be cycloid anteriorly or toward midline; body compressed, width at anus 11.9-28.1% (mean 15.2%) of SL; body stocky in appearance, body depth at anus 19.6-26.6%  of SL, body depth at first dorsal fin origin 14.3-27.7% of SL; head compressed, broader ventrally, slightly deeper than wide, HL 26.6-33.3% of SL; head depth at posterior preopercular margin 62.0-83.7% of HL; head width at posterior preopercular margin 53.7-82.2% of HL; head profile bluntly pointed; nape slightly convex behind eyes; mouth subterminal to nearly terminal, slightly oblique; jaws generally reaching to below anterior margin of eye; upper jaw length 32.0-41.5% of HL; upper lip smooth, narrower than lower, lower lip papillose close to teeth, with twist or fold posteriorly, forming triangular flap, lip interrupted at chin; eye moderate to relatively small in large specimens, dorsolateral, 22.7-30.5% of HL; preorbital width 18.2-27.8% of HL; snout bluntly pointed, 22.4-44.4% of HL; posterior naris round to almost triangular, close to anterior margin of eye; anterior naris in short tube, higher on posterior margin of eye, about level with middle of eye or somewhat ventral to it; interorbital narrow, 4.7-11.3% of HL; caudal peduncle compressed, length 12.9-18.6% (mean 16.1%) of SL; caudal peduncle depth 10.7-13.8% of SL (Ref. 92171).
Body shape (shape guide): fusiform / normal; Cross section: oval.				
				 
			
			
			
			
			
			
			
				Solitary or in loose groups in sand (Ref. 90102). Inhabits shallow coastal bays and estuaries on silty sandy substrates with sparse algae-reef or seagrasses (Ref. 48637).  Benthic (Ref. 58302). Feeds on organic matter and small invertebrates (Ref. 89972).			
			 
			
			
			
				
					Life cycle and mating behavior					
					Chín muồi sinh dục | Sự tái sinh sản | Đẻ trứng | Các trứng | Sự sinh sản | Ấu trùng				
				
				
				
				Benthic spawner.				
				
			
			 
				
				
				
					Randall, J.E. and D.W. Greenfield, 2001. A preliminary review of the Indo-Pacific gobiid fishes of the genus Gnatholepis. Ichthyological Bulletin of the J.L.B. Smith Institute of Ichthyology, 69:1-17. (Ref. 41653)
				
				 
			
			
			
							
					
						IUCN Red List Status   (Ref. 130435: Version 2025-1)
					
					
				 
					
			
			
			
			
			
			
			
				
					Threat to humans  
				
				
				
					  Harmless				
				
			 
			
			
			
			
				
					Human uses  				
				
				
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					Estimates based on models				
				
				
				Preferred temperature (Tài liệu tham khảo 
123201): 24.3 - 29, mean 27.7 °C (based on 808 cells).
				
				
					
					Phylogenetic diversity index  (Tài liệu tham khảo 
82804):  PD
50 = 0.5010   [Uniqueness, from 0.5 = low to 2.0 = high].					
													Bayesian length-weight: a=0.00977 (0.00444 - 0.02153), b=3.04 (2.85 - 3.23), in cm total length, based on LWR estimates for this (Sub)family-body shape (Ref. 
93245).
					
					Mức dinh dưỡng  (Tài liệu tham khảo 
69278):  3.4   ±0.5 se; based on size and trophs of closest relatives					
											
				
				
										
						Thích nghi nhanh  (Tài liệu tham khảo 
120179):  Chiêù cao, thời gian nhân đôi của chủng quần tối thiểu là dưới 15 tháng (Preliminary K or Fecundity.).					
											
				
									
					Fishing Vulnerability  (Ref. 
59153):  Low vulnerability (10 of 100). 
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						Nutrients  (Ref. 
124155):  Calcium = 133 [66, 296] mg/100g; Iron = 0.882 [0.415, 1.694] mg/100g; Protein = 18.2 [16.2, 20.0] %; Omega3 = 0.155 [0.068, 0.290] g/100g; Selenium = 23.3 [11.6, 57.7] μg/100g; VitaminA = 117 [29, 421] μg/100g; Zinc = 2.85 [1.88, 4.34] mg/100g (wet weight);