Environment: milieu / climate zone / depth range / distribution range
Marine; freshwater; brackish; benthopelagic; depth range - m (Ref. ), usually - m (Ref. ). Temperate; 0°C - ; 58°N - 34°N, 120°E - 161°E
Asia: Korean Peninsula, Japan, Kuril islands, Amur basin and Kamchatka Peninsula.
Size / Weight / Age
Maturity: Lm ?  range ? - ? cm
Max length : 6.5 cm SL male/unsexed; (Ref. 50519); 9.0 cm SL (female)
Length in freshwater is recorded at 6.5 cm TL; in the sea, it is recorded at 8.5-9.0 cm TL (Ref. 2058). Males build, guard and aerate the nest where the eggs are deposited (Ref. 205).
Life cycle and mating behavior
Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae
Males build, guard and aerate the nest where the eggs are deposited (Ref. 205).
Reshetnikov, Y.S., 2003. Atlas of Russian freshwater fishes, Vol. 2. Moscow Nauka. (Ref. 50519)
IUCN Red List Status (Ref. 124695)
Threat to humans
Estimates based on models
Phylogenetic diversity index (Ref. 82805
= 0.5020 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00457 (0.00223 - 0.00937), b=3.04 (2.85 - 3.23), in cm total length, based on LWR estimates for this species & (Sub)family-body (Ref. 93245
Trophic level (Ref. 69278
): 3.5 ±0.3 se; based on size and trophs of closest relatives
Resilience (Ref. 120179
): Medium, minimum population doubling time 1.4 - 4.4 years (Assuming Fec < 1,000).
Vulnerability (Ref. 59153
): Low vulnerability (21 of 100) .