Classification / Names
Common names | Synonyms | Catalog of Fishes(genus, species) | ITIS | CoL | WoRMS | Cloffa
Actinopteri (ray-finned fishes) > Gobiiformes
(Gobies) > Oxudercidae
(Mudskippers) > Periophthalminae
Etymology: Periophthalmus: Greek, peri = around + Greek, ophthalmos = eye (Ref. 45335).
Environment: milieu / climate zone / depth range / distribution range
Marine; brackish; demersal; depth range - m (Ref. ), usually - m (Ref. ). Tropical; 0°C -
Countries | FAO areas | Ecosystems | Occurrences | Point map | Introductions | Faunafri
Western Pacific: Thailand, Malaysia, Singapore and Indonesia.
Size / Weight / Age
Maturity: Lm ?  range ? - ? cm
Max length : 7.0 cm SL male/unsexed; (Ref. 46587)
Morphology | Morphometrics
(total): 9 - 12;
soft rays: 10 - 12. This species is characterized by the following: first dorsal fin pattern and coloration variable between individuals, typically, first dorsal fin rounded, moderate height, white margin, and one brown or black stripe inframarginally; the first spine of first dorsal fin always longest, rarely elongated; rounded to elliptical red-orange spots present ventral to inframarginal stripe, turning black in preservative; the second dorsal fin with yellow-orange margin and one black band inframarginally, rounded red-orange spots present on membrane between elements at base, turning black in preservative; live color with caudal and pectoral fins red-orange; prominent pelvic frenum, inner rays fused for less than half their length; ventero-posterior branchiostegal membrane dusky; D1 modally X; D2 modally I,12; A I,11; caudal fin modally 14 segmented rays; 48-60 lateral scales; 17-22 predorsal scales (Ref. 83659).
Life cycle and mating behavior
Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae
Jaafar, Z., M. Perrig and L.M. Chou, 2009. Periophthalmus variabilis (Teleostei: Gobiidae: Oxudercinae), a valid species of mudskipper, and a re-diagnosis of Periophthalmus novemradiatus. Zool. Sci. 26:309-314. (Ref. 83659)
IUCN Red List Status (Ref. 124695)
Threat to humans
CountriesFAO areasEcosystemsOccurrencesIntroductionsStocksEcologyDietFood itemsFood consumptionRation
Common namesSynonymsMetabolismPredatorsEcotoxicologyReproductionMaturitySpawningSpawning aggregationFecundityEggsEgg development
ReferencesAquacultureAquaculture profileStrainsGeneticsAllele frequenciesHeritabilityDiseasesProcessingNutrientsMass conversion
Estimates based on models
Phylogenetic diversity index (Ref. 82805
= 0.5000 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00977 (0.00571 - 0.01672), b=2.91 (2.76 - 3.06), in cm total length, based on LWR estimates for this species & Genus-body shape (Ref. 93245
Trophic level (Ref. 69278
): 3.2 ±0.4 se; based on size and trophs of closest relatives
Vulnerability (Ref. 59153
): Low vulnerability (10 of 100) .