Environment: milieu / climate zone / depth range / distribution range
Marine; demersal; depth range - m (Ref. ), usually - m (Ref. ). Tropical; 0°C -
Northwest Pacific: Japan (from Kanagawa Prefecture and the Goto Islands southward), Taiwan, and the South China Sea.
Size / Weight / Age
Maturity: Lm ?  range ? - ? cm
Max length : 8.0 cm TL male/unsexed; (Ref. 637)
Occurs solitarily around boulders scattered on sandy and muddy bottoms in bays. Breeding occurs from July to September. Males brood eggs in the mouth (Ref. 559).
Life cycle and mating behavior
Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae
Mouthbrooders (Ref. 240). Distinct pairing during courtship and spawning (Ref. 205).
Masuda, H., K. Amaoka, C. Araga, T. Uyeno and T. Yoshino, 1984. The fishes of the Japanese Archipelago. Vol. 1. Tokai University Press, Tokyo, Japan. 437 p. (text). (Ref. 559)
IUCN Red List Status (Ref. 124695)
Threat to humans
ReferencesAquacultureAquaculture profileStrainsGeneticsAllele frequenciesHeritabilityDiseasesProcessingNutrientsMass conversion
Estimates based on models
Preferred temperature (Ref. 115969
): 20.9 - 29.2, mean 28.7 °C (based on 1292 cells).
Phylogenetic diversity index (Ref. 82805
= 0.5000 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.01479 (0.00690 - 0.03171), b=3.09 (2.91 - 3.27), in cm total length, based on LWR estimates for this (Sub)family-body shape (Ref. 93245
Trophic level (Ref. 69278
): 3.4 ±0.5 se; based on size and trophs of closest relatives
Resilience (Ref. 120179
): High, minimum population doubling time less than 15 months (Preliminary K or Fecundity.).
Vulnerability (Ref. 59153
): Low vulnerability (10 of 100) .