Teleostei (teleosts) > 
Blenniiformes (Blennies) > 
Tripterygiidae (Triplefin blennies) > Tripterygiinae						
							
							Etymology: Helcogramma: Greek, helkos, -eos, -ous = ulcer, sore + Greek, gramma = letter, mark (Ref. 45335).
More on author: Fowler.						
					
				
					Environment: milieu / climate zone / depth range / distribution range					
						Ecology					
				
				
				
					Marine;  demersal; depth range 0 - 8 m (Ref. 90102). Subtropical				
				 
			
			
			
				
				
				
					Western Pacific:  Japan, Ryukyu Islands, Taiwan, Hong Kong, Philippines, Viet Nam, Thailand, Malaysia, Indonesia, and Vanuatu.
				
				
			
			
				
					Size / Weight / Age
				
				
				
					Maturity: Lm ?  range ? - ? cm
 Max length : 3.2 cm SL male/unsexed; (Ref. 27223)				
				 
			
			
						
				
					Short description					
					Identification keys | Morphology | Morphometrics					
				
				
			
			
			
			
			
			
			
				Adults are found at depths of 8 m and shallower, in surge channels and exposed rocky and coralline boulders (Ref. 90102).  Eggs are hemispherical and covered with numerous sticky threads that anchor them in the algae on the nesting sites (Ref. 240).  Larvae are planktonic which occur primarily in shallow, nearshore waters (Ref. 94114).			
			 
			
			
			
				
					Life cycle and mating behavior					
					Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae				
				
				
				
								
				
			
			 
				
				
				
					Fricke, R., 1997. Tripterygiid fishes of the western and central Pacific, with descriptions of 15 new species, including an annotated checklist of world Tripterygiidae (Teleostei). Theses Zool. 29:1-607. (Ref. 27223)
				
				 
			
			
			
							
					
						IUCN Red List Status   (Ref. 130435: Version 2025-1)
					
					
				 
					
			
			
			
			
			
			
			
				
					Threat to humans  
				
				
				
					  Harmless				
				
			 
			
			
			
			
				
					Human uses  				
				
				
									
				
				
			
			
						
			
			
			
				
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					Estimates based on models				
				
				
				Preferred temperature (Ref. 
123201): 24.9 - 29.3, mean 28.6 °C (based on 1707 cells).
				
				
					
					Phylogenetic diversity index  (Ref. 
82804):  PD
50 = 0.5000   [Uniqueness, from 0.5 = low to 2.0 = high].					
													Bayesian length-weight: a=0.00617 (0.00288 - 0.01322), b=3.04 (2.86 - 3.22), in cm total length, based on LWR estimates for this (Sub)family-body shape (Ref. 
93245).
					
					Trophic level  (Ref. 
69278):  3.0   ±0.3 se; based on size and trophs of closest relatives					
											
				
				
										
						Resilience  (Ref. 
120179):  High, minimum population doubling time less than 15 months (Preliminary K or Fecundity.).					
											
				
									
					Fishing Vulnerability  (Ref. 
59153):  Low vulnerability (10 of 100). 
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