Environment: milieu / climate zone / depth range / distribution range
Freshwater; benthopelagic; depth range - m (Ref. ), usually - m (Ref. ). Temperate; 0°C - ; 55°N - 42°N, 141°E - 145°E
Asia: Hokkaido, Japan and Sakhalin, Russian Federation.
Size / Weight / Age
Maturity: Lm ?  range ? - ? cm
Max length : 7.0 cm TL male/unsexed; (Ref. 2058)
Adults inhabit weedy streams. Males build, guard and aerate the nest where the eggs are deposited just above the bottom (Ref. 205, 50519).
Life cycle and mating behavior
Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae
Males build, guard and aerate the nest where the eggs are deposited (Ref. 205).
Reshetnikov, Y.S., 2003. Atlas of Russian freshwater fishes, Vol. 2. Moscow Nauka. (Ref. 50519)
IUCN Red List Status (Ref. 124695)
Threat to humans
ReferencesAquacultureAquaculture profileStrainsGeneticsAllele frequenciesHeritabilityDiseasesProcessingNutrientsMass conversion
Estimates based on models
Phylogenetic diversity index (Ref. 82805
= 0.5020 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00437 (0.00181 - 0.01052), b=3.06 (2.85 - 3.27), in cm total length, based on LWR estimates for this (Sub)family-body shape (Ref. 93245
Trophic level (Ref. 69278
): 3.4 ±0.2 se; based on size and trophs of closest relatives
Resilience (Ref. 120179
): Medium, minimum population doubling time 1.4 - 4.4 years (Assuming Fec < 1,000).
Vulnerability (Ref. 59153
): Low vulnerability (13 of 100) .