This species have the following characteristics: pyloric caeca usually 4-8 in 2 whorls, typically 2-3 in first 2 whorls and 1in third; usually 3 rudimentary gill rakers on the upper arm of the first arch and 5-7 developed rakers on lower arm, 1-6 rudimentary rakers anteriorly on the lower arm; total gill rakers 9-14; dorsal-fin origin between neural spines 3-4; roof of mouth posterior to the palatine and vomerine tooth patches blackish, also the gill bars, upper pharyngeal region, esophagus, caeca, and anterior part of the intestine; pectoral fins dusky; dorsal and anal fins have a dark edge; males have the anterior rib thickened (Ref. 91765).