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|Sound produced||yes, active sound production|
|Type of sound produced||thumps & knocks|
|Sound production organ||pharyngeal teeth & swim bladder|
|Sound mechanism||pharyngeal teeth stridulation augmented by adjacent swim bladder|
|Behavioural context||spontaneous sounds during competitive feeding|
|Reference||Fish, M.P. and W.H. Mowbray, 1970|
|Remark||recording made in outdoor pen with bandpass filter: 30 - 1200 Hz, high pass noise reduction filter applied to recording & amplified by 3 dB|