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Click here to download the sound file.| Sound produced | yes, active sound production |
| Type of sound produced | knocks, thumps, escape sounds |
| Sound production organ | swim bladder |
| Sound mechanism | vibration by contraction of associated muscles |
| Behavioural context | under duress (handling & electric stimulation) |
| Reference | Fish, M.P. and W.H. Mowbray, 1970 |
| Remark | recorded with highpass filter: 70 Hz |