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| Sound produced | yes, active sound production |
| Type of sound produced | deep booms, sustained rumbles, grunts, scrapes |
| Sound production organ | swim bladder firmly encased in peritoneum, teeth |
| Sound mechanism | vibration by contraction of associated axial muscles & bilateral muscle behind opercles, scraping of teeth |
| Behavioural context | booms & rumbles: spontaneous & with mild annoyance, loud grunts under duress, grunts & scraping during competitive feeding |
| Reference | Fish, M.P. and W.H. Mowbray, 1970 |
| Remark |