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Click here to download the sound file.| Sound produced | yes, active sound production |
| Type of sound produced | grunts |
| Sound production organ | swim bladder |
| Sound mechanism | vibration by contraction of the drumming muscles associcated with the swimbladder |
| Behavioural context | courtship sound presumably by males during spawning period |
| Reference | Finstad, J.L. and J.T. Nordeide, 2001 |
| Remark | recording was carried out in land-based fiberglass tanks during spawning period |