List of Nominal Species of Mullidae
(Goatfishes)
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Nominal Species (n = 152) Present allocation (n = 88) Ref.
Parupeneus angulatus Randall & Heemstra, 2009 Parupeneus angulatus 82838
Upeneus arge Jordan & Evermann, 1903 Upeneus taeniopterus 83903
Mullus argentinae Hubbs & Marini, 1933 Mullus argentinae 12736
Pennon armatoides Whitley, 1955 Upeneus sundaicus 75154
Upeneus asymmetricus (non Lachner, 1954) Upeneus australiae 98481
Upeneus asymmetricus Lachner, 1954 Upeneus asymmetricus 3132
Upeneus atrocingulatus Kner, 1870 Parupeneus multifasciatus 54393
Pseudupeneus aurantiacus Seale, 1906 Parupeneus cyclostomus 54393
Parupeneus aurantius Fourmanoir, 1966 Parupeneus biaculeatus 54393
Mullus auriflamma Forsskål, 1775 Parupeneus forsskali 54393
Upeneus australiae Kim & Nakaya, 2002 Upeneus australiae 43139
Mulloidichthys ayliffe Uiblein, 2011 Mulloidichthys ayliffe 86817
Mullus barbatus Linnaeus, 1758 Mullus barbatus barbatus 3397
Mullus barbatus auratus Jordan & Gilbert, 1882 Mullus auratus 9626
Mullus barbatus ponticus Essipov, 1927 Mullus barbatus ponticus 4845
Upeneus barberinoides Bleeker, 1852 Parupeneus barberinoides 54393
Mullus barberinus Lacepède, 1801 Parupeneus barberinus 54393
Mullus bensasi Temminck & Schlegel, 1843 Upeneus japonicus 10687
Upeneus biaculeatus Richardson, 1846 Parupeneus biaculeatus 54393
Mullus bifasciatus Lacepède, 1801 Parupeneus trifasciatus 45589
Upeneus bilineatus Valenciennes, 1831 Upeneus sulphureus 94476
Upeneus brandesii Bleeker, 1851 Parupeneus pleurostigma 54393
Mullus chryserydros Lacepède, 1801 Parupeneus cyclostomus 54393
Upeneus chryserythrus Günther, 1873 Parupeneus cyclostomus 54393
Pseudupeneus chrysonemus Jordan & Evermann, 1903 Parupeneus chrysonemus 26282
Mullus chrysopleuron Temminck & Schlegel, 1843 Parupeneus chrysopleuron 54393
Sciaena ciliata Lacepède, 1802 Parupeneus ciliatus 54393
Upeneus cinnabarinus Cuvier, 1829 Parupeneus heptacanthus 54393
Atahua clarki Phillipps, 1941 Upeneichthys lineatus 88002
Upeneus crassilabris Valenciennes, 1831 Parupeneus crassilabris 54393
Upeneus crassilabris (non Valenciennes, 1831) Parupeneus insularis 45589
Upeneus crosnieri Fourmanoir & Guézé, 1967 Upeneus guttatus 83903
Mullus cyclostomus Lacepède, 1801 Parupeneus cyclostomus 54393
Upeneus cyprinoides Valenciennes, 1831 Parupeneus ciliatus 54393
Upeneus davidaromi Golani, 2001 Upeneus davidaromi 44136
Upeneus dentatus Gill, 1862 Mulloidichthys dentatus 12965
Parupeneus diagonalis Randall, 2004 Parupeneus diagonalis 54393
Mullus dispilurus Playfair, 1867 Parupeneus rubescens 50838
Upeneoides doriae Günther, 1869 Upeneus doriae 13710
Mullus dubius Temminck & Schlegel, 1843 Parupeneus chrysopleuron 54393
Mulloides erythrinus Klunzinger, 1884 Mulloidichthys vanicolensis 2872
Pseudupeneus eutaeniatus Fowler, 1944 Parupeneus ciliatus 54393
Upeneoides fasciolatus Day, 1868 Upeneus moluccensis 2795
Upeneoides filifer Ogilby, 1910 Upeneus filifer 46206
Mullus flavolineatus Lacepède, 1801 Mulloidichthys flavolineatus 5405
Pseudupeneus forsskali Fourmanoir & Guézé, 1976 Parupeneus forsskali 54393
Upeneus francisi Randall & Guézé, 1992 Upeneus francisi 10665
Parupeneus fraserorum Randall & King, 2009 Parupeneus fraserorum 82839
Upeneus fraterculus Valenciennes, 1831 Parupeneus ciliatus 54393
Mullus fuscatus Rafinesque, 1810 Mullus surmuletus 7313
Upeneus grandisquamis Gill, 1863 Pseudupeneus grandisquamis 12965
Upeneus griseofrenatus Kner, 1868 Parupeneus indicus 54393
Upeneoides guttatus Day, 1868 Upeneus guttatus 54621
Upeneus heemstra Uiblein & Gouws, 2014 Upeneus heemstra 95469
Sciaena heptacantha Lacepède, 1802 Parupeneus heptacanthus 54393
Upeneus immaculatus Bennett, 1831 Parupeneus cyclostomus 54393
Mullus indicus Shaw, 1803 Parupeneus indicus 54393
Upeneus indicus Uiblein & Heemstra, 2010 Upeneus indicus 83903
Parupeneus insularis Randall & Myers, 2002 Parupeneus insularis 45589
Pseudupeneus ischyrus Snyder, 1907 Parupeneus ciliatus 54393
Upeneus itoui Yamashita, Golani & Motomura, 2011 Upeneus itoui 88711
Upeneus jansenii Bleeker, 1856 Parupeneus jansenii 54393
Mullus japonicus Houttuyn, 1782 Upeneus japonicus 10687
Pseudupeneus jeffi Ogilby, 1908 Parupeneus spilurus 75154
Parupeneus kneri Bleeker, 1872 Parupeneus biaculeatus 54393
Mullus lateristriga Cuvier, 1829 Parupeneus macronemus 54393
Upeneus lombok Uiblein & White, 2015 Upeneus lombok 102936
Parupeneus louise Randall, 2004 Parupeneus louise 54393
Upeneus luteus Valenciennes, 1831 Parupeneus cyclostomus 54393
Upeneus luzonius Jordan & Seale, 1907 Upeneus sundaicus 9947
Upeneus luzonius (non Jordan & Seale, 1907) Upeneus tragula 95469
Upeneus luzonius Jordan & Seale, 1907 Upeneus luzonius 8631
Mullus macronemus Lacepède, 1801 Parupeneus macronemus 54393
Mullus maculatus Bloch, 1793 Pseudupeneus maculatus 9626
Upeneus malabaricus Cuvier, 1829 Parupeneus indicus 54393
Upeneus margarethae Uiblein & Heemstra, 2010 Upeneus margarethae 83903
Parupeneus margaritatus Randall & Guézé, 1984 Parupeneus margaritatus 54393
Upeneus martinicus Cuvier, 1829 Mulloidichthys martinicus 3792
Upeneus mascareinsis Fourmanoir & Guézé, 1967 Upeneus mascareinsis 33390
Mullus microps Bliss, 1883 Parupeneus cyclostomus 54393
Mulloidichthys mimicus Randall & Guézé, 1980 Mulloidichthys mimicus 9710
Mulloidichthys mimicus (non Randall & Guézé, 1980) Mulloidichthys ayliffe 86817
Parupeneus minys Randall & Heemstra, 2009 Parupeneus minys 82838
Pseudupeneus moana Jordan & Seale, 1906 Parupeneus multifasciatus 54393
Parupeneus moffitti Randall & Myers, 1993 Parupeneus moffitti 10673
Upeneoides moluccensis Bleeker, 1855 Upeneus moluccensis 2795
Upeneus mouthami Randall & Kulbicki, 2006 Upeneus mouthami 58005
Mullus multifasciatus Quoy & Gaimard, 1825 Parupeneus multifasciatus 54393
Parupeneus nansen Randall & Heemstra, 2009 Parupeneus nansen 82838
Upeneus niebuhri Guézé, 1976 Upeneus niebuhri 95469
Upeneus nigromarginatus Bos, 2014 Upeneus nigromarginatus 97569
Upeneus oligospilus Lachner, 1954 Upeneus oligospilus 83903
Upeneus oligospilus (non Lachner, 1954) Upeneus heemstra 95469
Pseudupeneus orientalis Fowler, 1933 Parupeneus orientalis 54393
Upeneus oxycephalus Bleeker, 1856 Parupeneus cyclostomus 54393
Upeneus parvus Poey, 1852 Upeneus parvus 9626
Mulloides pfluegeri Steindachner, 1900 Mulloidichthys pfluegeri 12965
Upeneus pleurospilos Bleeker, 1853 Parupeneus heptacanthus 54621
Upeneus pleurostigma Bennett, 1831 Parupeneus pleurostigma 54393
Mullus pleurotaenia Playfair, 1867 Parupeneus ciliatus 54393
Upeneus pori Ben-Tuvia & Golani, 1989 Upeneus pori 11441
Upeneus porosus Cuvier, 1829 Upeneichthys lineatus 50838
Pseudupeneus porphyreus (non Jenkins, 1903) Parupeneus ciliatus 55073
Pseudupeneus porphyreus Jenkins, 1903 Parupeneus porphyreus 54393
Parupeneus posteli Fourmanoir & Guézé, 1967 Parupeneus posteli 33390
Upeneus prayensis Cuvier, 1829 Pseudupeneus prayensis 7313
Parupeneus procerigena Kim & Amaoka, 2001 Parupeneus procerigena 45339
Upeneus quadrilineatus Cheng & Wang, 1963 Upeneus quadrilineatus 559
Mullus radiatus Shaw, 1803 Parupeneus cyclostomus 54393
Upeneus randalli Uiblein & Heemstra, 2011 Upeneus randalli 88844
Mullus ruber Lacepède, 1801 Mullus barbatus barbatus 7313
Mullus rubescens Lacepède, 1801 Parupeneus rubescens 5405
Mullus russelii Cuvier, 1829 Parupeneus indicus 54393
Upeneus saffordi Seale, 1901 Parupeneus cyclostomus 54393
Upeneus saiab Uiblein & Lisher, 2013 Upeneus saiab 94148
Mulloides samoensis Günther, 1874 Mulloidichthys flavolineatus 2334
Upeneus semifasciatus Macleay, 1883 Parupeneus crassilabris 54393
Pseudupeneus seychellensis Smith & Smith, 1963 Parupeneus seychellensis 82838
Pseudupeneus seychellensis (non Smith & Smith, 1963) Parupeneus heptacanthus 82838
Upeneus seychellensis Uiblein & Heemstra, 2011 Upeneus seychellensis 87809
Upeneus signatus Günther, 1867 Parupeneus spilurus 75154
Upeneus spilurus Bleeker, 1854 Parupeneus spilurus 26282
Upeneus stenopsis Uiblein & McGrouther, 2012 Upeneus stenopsis 92287
Upeneichthys stotti Hutchins, 1990 Upeneichthys stotti 38963
Upeneus suahelicus Uiblein & Heemstra, 2010 Upeneus suahelicus 83903
Mullus subvittatus Temminck & Schlegel, 1843 Upeneus subvittatus 26282
Parupeneus sufflavus Whitley, 1941 Parupeneus ciliatus 54393
Upeneus sulphureus (non Cuvier, 1829) Upeneus doriae 12965
Upeneus sulphureus Cuvier, 1829 Upeneus sulphureus 5405
Upeneoides sundaicus Bleeker, 1855 Upeneus sundaicus 4899
Upeneus supravittatus Uiblein & Heemstra, 2010 Upeneus supravittatus 83903
Mullus surmuletus Linnaeus, 1758 Mullus surmuletus 7313
Mullus surmuletus lineatus Bloch & Schneider, 1801 Upeneichthys lineatus 26282
Upeneus taeniatus Kner, 1865 Parupeneus biaculeatus 54393
Upeneus taeniatus (non Kner, 1865) Parupeneus ciliatus 54393
Upeneus taeniopterus Cuvier, 1829 Upeneus taeniopterus 83903
Upeneoides tokisensis Döderlein, 1883 Upeneus japonicus 10687
Upeneus torres Uiblein & Gledhill, 2014 Upeneus torres 98481
Upeneus tragula (non Richardson, 1846) Upeneus oligospilus 83903
Upeneus tragula (non Richardson, 1846) Upeneus niebuhri 95469
Upeneus tragula Richardson, 1846 Upeneus tragula 1602
Upeneus tragula (non Richardson, 1846) Upeneus heemstra 95469
Mullus trifasciatus Lacepède, 1801 Parupeneus trifasciatus 280
Upeneus vanicolensis Valenciennes, 1831 Mulloidichthys vanicolensis 5405
Upeneus vanuatu Uiblein & Causse, 2013 Upeneus vanuatu 93585
Upeneus velifer Smith & Swain, 1882 Parupeneus multifasciatus 54393
Mullus vittatus Forsskål, 1775 Upeneus vittatus 5405
Upeneus vlamingii Cuvier, 1829 Upeneichthys vlamingii 33839
Upeneus waigiensis Cuvier, 1829 Parupeneus indicus 54393
Upeneus xanthogrammus Gilbert, 1892 Upeneus xanthogrammus 11035
Pseudupeneus xanthopurpureus Fourmanoir, 1957 Parupeneus heptacanthus 54393
Parupeneus xanthospilurus Bleeker, 1875 Parupeneus cyclostomus 54393
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cfm script by eagbayani, 10.5.01 ,  last modified by cmilitante, 29/11/12