| 主要参考文献 | Zander, C.D., 1986 |
|---|---|
| 附注 | Occurs in the intertidal zone and shallow bottoms, on rocks or sand between pebbles and vegetation. Often in brackish waters down to 5 ppt. Inhabits crevices or piddock holes, males remain in cavities above water-level during low tide. Feeds on benthic invertebrates, mainly molluscs, also algae. |
| Marine - Neritic | Marine - Oceanic | Brackishwater | Freshwater | |
|---|---|---|---|---|
| Marine zones / Brackish and freshwater bodies |
|
|
|
|
| Substrate | : rocky; |
|---|---|
| Substrate Ref. | |
| Special habitats | |
| Special habitats Ref. |
| 参考文献 | |
|---|---|
| Associations | |
| Associated with | |
| Association remarks | |
| Parasitism |
| 摄取食物的类型 | plants/detritus+animals (troph. 2.2-2.79) |
|---|---|
| Feeding type Ref. | Zander, C.D., 1986 |
| 摄取食物的习惯 | variable |
| Feeding habit Ref. | Zander, C.D., 1986 |
| Estimation method | 最初的样本 | 不捕获族群 | 附注 | ||
|---|---|---|---|---|---|
| Troph | s.e. | Troph | s.e. | ||
| 从食物内容 | 3.59 | 2.96 | 0.17 | Troph of adults from 1 study. | |
| 从个别的食物项目 | 3.02 | 0.34 | Trophic level estimated from a number of food items using a randomized resampling routine. | ||
| 参考文献 | Goldschmid, A., K. Kotrschal and P. Wirtz, 1984 | ||||