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Alosa immaculata  Bennett, 1835

Pontic shad
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Alosa immaculata   AquaMaps   Data sources: GBIF OBIS
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Image of Alosa immaculata (Pontic shad)
Alosa immaculata
Picture by Kutsarov, Y.

Classification / Names Common names | Synonyms | Catalog of Fishes (gen., sp.) | ITIS | CoL | WoRMS | Cloffa

Actinopterygii (ray-finned fishes) > Clupeiformes (Herrings) > Clupeidae (Herrings, shads, sardines, menhadens) > Alosinae
Etymology: Alosa: Latin, alausa = a fish cited by Ausonius and Latin, halec = pickle, dealing with the Greek word hals = salt; it is also the old Saxon name for shad = "alli" ; 1591 (Ref. 45335).   More on author: Bennett.

Environment / Climate / Range Ecology

Marine; freshwater; brackish; pelagic-neritic; anadromous (Ref. 51243); depth range 3 - 90 m (Ref. 10439).   Temperate; 50°N - 41°N, 27°E - 44°E

Size / Weight / Age

Maturity: Lm ?  range ? - 23.4 cm
Max length : 39.0 cm SL male/unsexed; (Ref. 188); max. reported age: 7 years (Ref. 10439)

Short description Morphology | Morphometrics

Dorsal spines (total): 0; Anal spines: 0. Body fairly elongate, more `herring-like' than `shad-like'. Gill rakers rather thin, usually equal to or a little shorter than gill filaments. Teeth well developed in both jaws. Resembles A. caspia, which usually has more Gill rakers (50 to 180, much longer than gill filaments), poorly developed teeth and a deeper, `shad-like' body; A. maeotica has fewer Gill rakers (33 to 36).

Distribution Countries | FAO areas | Ecosystems | Occurrences | Point map | Introductions | Faunafri

Eurasia: Black Sea and Sea of Azov ( in sea and in the Don, Danube and other rivers, as much as 567 km up the Don and as far as Kiev on the Dneiper before the dam was built). Recognized sub-species (Ref. 683): Alosa pontica pontica in the Black Sea and rivers feeding it; Alosa pontica kessleri and Alosa pontica volgensis in the Caspian Sea. Appendix III of the Bern Convention (protected fauna).

Biology     Glossary (e.g. epibenthic)

Thai species is pelagic at sea, in deep water. It migrates to middle reaches of large rivers, spawning where current is strongest, close to the surface, often at 2-3 m depth in the main channel. Migrates upriver to spawn at 3 years, rarely earlier and only a few individuals spawn two seasons. It appears along the coast in March to April, enters rivers when temperatures reach about 6-9°C, between late March and late April; migration usually peaks in May. Spawning starts when temperature rises above 15°C in April to August; usually between 1 and 8 p.m. Pelagic eggs. Spent individuals return to the sea to feed. Juveniles inhabit floodplain and shallow riverine habitats, migrate to the sea or estuarine habitats during first summer; in autumn return to the sea until maturity. When at sea, it feeds on a wide variety of zooplankton (mainly crustaceans, (Crangon, Upogebia, Idothea, gammarids) and small fish (Engraulis, Clupeonella, Sprattus). Reduction of spawning sites and migration routes are caused by impoundment of main rivers. Heavy over fishing apparently reduced all populations during the first decade of the 20th century (Ref. 59043). Reaches a smaller size (30 cm SL, usually 14-18) in Don River; a smaller form reaches only 21 cm.

Life cycle and mating behavior Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae

Eggs are pelagic(Ref. 59043).

Main reference Upload your references | References | Coordinator | Collaborators

Kottelat, M., 1997. European freshwater fishes. Biologia 52, Suppl. 5:1-271. (Ref. 13696)

IUCN Red List Status (Ref. 96402)

  Vulnerable (VU) (B2ab(v))

CITES (Ref. 94142)

Not Evaluated

Threat to humans

  Harmless




Human uses

Fisheries: minor commercial
FAO(fisheries: production; publication : search) | FisheriesWiki | Sea Around Us

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Internet sources

Alien/Invasive Species database | BHL | Cloffa | Websites from users | Check FishWatcher | CISTI | Catalog of Fishes (gen., sp.) | DiscoverLife | Faunafri | Fishtrace | GenBank(genome, nucleotide) | GOBASE | Google Books | Google Scholar | Google | IGFA World Record | iSpecies | National databases | PubMed | Scirus | SeaLifeBase | Tree of Life | uBio | Wikipedia(Go, Search) | World Records Freshwater Fishing | Zoobank | Zoological Record

Estimates of some properties based on models

Phylogenetic diversity index (Ref. 82805):  PD50 = 0.5000   [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.01349 (0.00785 - 0.02318), b=2.99 (2.84 - 3.14), based on LWR estimates for species & Genus-body shape (Ref. 93245).
Trophic Level (Ref. 69278):  3.9   ±0.67 se; Based on food items.
Resilience (Ref. 69278):  Medium, minimum population doubling time 1.4 - 4.4 years (K=0.32; tm=2; tmax=9).
Vulnerability (Ref. 59153):  Low to moderate vulnerability (35 of 100) .
Price category (Ref. 80766):   High.