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Alosa fallax  (Lacepède, 1803)

Twaite shad
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Alosa fallax
Picture by Stergiou, K.I.

Classification / Names Common names | Synonyms | Catalog of Fishes (gen., sp.) | ITIS | CoL | WoRMS | Cloffa

Actinopterygii (ray-finned fishes) > Clupeiformes (Herrings) > Clupeidae (Herrings, shads, sardines, menhadens) > Alosinae
Etymology: Alosa: Latin, alausa = a fish cited by Ausonius and Latin, halec = pickle, dealing with the Greek word hals = salt; it is also the old Saxon name for shad = "alli" ; 1591 (Ref. 45335).

Issue
5 subspecies known.

Environment / Climate / Range Ecology

Marine; freshwater; brackish; pelagic-neritic; anadromous (Ref. 51243); depth range ? - 300 m (Ref. 10541).   Temperate; 70°N - 27°N, 25°W - 42°E

Length at first maturity / Size / Weight / Age

Maturity: Lm 32.5  range ? - ? cm
Max length : 60.0 cm SL male/unsexed; (Ref. 35388); common length : 40.0 cm SL male/unsexed; (Ref. 2945); max. published weight: 1.5 kg (Ref. 188); max. reported age: 25 years (Ref. 556)

Short description Morphology | Morphometrics

Dorsal spines (total): 4 - 6; Dorsal soft rays (total): 12-16; Anal spines: 3-4; Anal soft rays: 16 - 22; Vertebrae: 55 - 59. Body slightly compressed, fairly deep with scutes along belly. Upper jaw notched, lower jaw fitting into it (Ref. 188). 31-50 (rarely up to 60) thick gill rakers on the lower arc (Ref. 59043). Large, thin scales (Ref. 51442). Deep blue dorsally, becoming greenish brown or golden on the sides and silvery ventrally. A dark spot posterior to gill opening followed by 6-10 similar spots (sometimes faint or absent) along flank (Refs. 188, 88187). Also Ref. 2196.

Distribution Countries | FAO areas | Ecosystems | Occurrences | Point map | Introductions | Faunafri

Northeast Atlantic: from the British Isles and southern Norway to Morocco, including the Baltic, Mediterranean and Black Seas (Refs. 188, 26334, 51442)). Several subspecies have been recognized based on the number of gill rakers and geographical location (Ref. 10541) and some have since been given species-status (Ref. 59043). Listed in Appendix III of the Bern Convention (2002). Listed in Annex II and V of the EC Habitats Directive (2007).

Biology     Glossary (e.g. epibenthic)

Amphihaline species (Ref. 51442), schooling and strongly migratory (Ref. 188). Adults are usually found in open waters along the coast (Refs. 59043, 89486); juveniles are usually found along estuaries and near the shore (Ref. 59043), possibly making vertical diurnal movements synchronized with the tides; they remain in estuaries for over one year (Ref. 89630). Migrates to major rivers to spawn; also reported to spawn in small rivers. Several landlocked (lake) non-migratory populations exist (Ref. 10541). Ichthyophagous, feeds on small fishes and crustaceans, the young taking the fry of herrings, sprats and gobies (Ref. 188, Ref. 51442). Females grow faster and are always larger than males of the same age (Ref. 10541). Very locally distributed due to pollution and impoundment of large rivers throughout Europe and most populations declined during the first decade of the 20th century, but seem to have stabilized at a low level since then (Ref. 59043). It has been suggested that members of the genus Alosa are hearing specialists with the American shad (Alosa sapidissima) having been found to detect and respond to sounds up to at least 180 kHz (Ref. 89631). This may aid in predator avoidance (e.g. cetaceans) (Ref. 89632). Hybridization between this species and the allis shad (Alosa alosa) has been reported from the Rhine (Ref. 89633) as well as rivers in France and Algeria (Ref. 10541). There is some evidence that indicates that shad hybrids may reproduce (Ref. 27567).

Life cycle and mating behavior Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae

Adults in the sea begin to congregate near mouths of estuaries in April. Enter estuaries and ascend rivers in May and June when water temperature is between 10-14 °C (Refs. 188, 51442, 59043, 89636). Males begin such movements at 2-3 years, females at 3-4 years (Ref. 59043). Although movement upstream is usually limited to a few kilometres above the brackish zone (Refs. 59043, 89486), spawning has also been reported in non-tidal freshwater areas up to 400 km upstream (Ref. 89637). Gametogenesis occurs in the estuaries. Early arrivals in the rivers are mostly males, with the sex ratio becoming more equal with the later arrivals (Ref. 42360). Spawning movements occur with spring tides and peak when river discharge levels are high (Refs. 89636, 89638). However, when flows are too high, movements upstream become limited (Refs. 89636, 89639). Spawn when water temperature is anywhere between 12-22 °C (Ref. 10541). Move to riverine spawning grounds at night; spawn in large, very noisy schools near surface and leave these areas before daybreak (Ref. 10541). Spawning sites consist of sand and gravel areas with flowing water (Ref. 10541). Spent adults return to the sea and may spawn for 3-4 seasons throughout their lifetime (Refs. 30578, 51442, 59043). Most individuals will have lost 22 % of their body weight after spawning (Ref. 89640). There is some evidence that most individuals return to their natal rivers to spawn (Refs. 10541, 59043). Eggs either drift with the current or sink to the bottom (Ref. 59043, 89641). Eggs hatch after 2-8 days, depending on water temperature (optimal 15-25 °C) (Refs. 35387, 41851). Larvae and juveniles move towards the estuaries and river mouths during their first summer and to the sea at the end of their second year (Ref. 59043). Males mature mainly between the ages of 2-5 years, females between 3-7 years (Ref. 188, 2163, 10541). Length at maturity is between 30-40 cm total length (Ref. 88187).

Main reference Upload your references | References | Coordinator | Collaborators

Whitehead, P.J.P., 1985. FAO Species Catalogue. Vol. 7. Clupeoid fishes of the world (suborder Clupeioidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies and wolf-herrings. FAO Fish. Synop. 125(7/1):1-303. Rome: FAO. (Ref. 188)

IUCN Red List Status (Ref. 96402)

CITES (Ref. 94142)

Not Evaluated

Threat to humans

  Harmless




Human uses

Fisheries: minor commercial
FAO(fisheries: production; publication : search) | FisheriesWiki | Sea Around Us

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Estimates of some properties based on models

Phylogenetic diversity index (Ref. 82805):  PD50 = 0.5000   [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00617 (0.00507 - 0.00749), b=3.02 (2.97 - 3.07), based on LWR estimates for this species (Ref. 93245).
Trophic Level (Ref. 69278):  4.0   ±0.4 se; Based on diet studies.
Resilience (Ref. 69278):  Medium, minimum population doubling time 1.4 - 4.4 years (K=0.21-0.38; tm=2-7; tmax=25; Fec>10,000).
Vulnerability (Ref. 59153):  Moderate to high vulnerability (50 of 100) .
Price category (Ref. 80766):   Low.