| Main Ref. | Harada, Y., S. Harada, I. Kinoshita, M. Tanaka and M. Tagawa, 2003 |
| Mode | dioecism |
| Fertilization | external |
| Mating type | |
| Spawning frequency | |
| Spawning aggregation | Ref. |
| Batch spawner | Ref. |
| Reproductive guild |
guarders nesters |
| Parental Care | paternal |
| Description of life cycle and mating behavior | Females spawn in nests under stones (Ref. 101248). The egg clusters are cared for by a male until hatching (Ref. 101248). |
| Search for more references on reproduction | Scirus |